English: Model of two adjacent salmon protamine molecules (blue) wrapped around and bound within the major groove the DNA helix (white). Protamine binding neutralizes the phosphodiester backbone of DNA, causing it to coil into toroidal structures.
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Uploaded a work by Rod Balhorn from [https://doi.org/10.1186/gb-2007-8-9-227 Balhorn, R. (2007). The protamine family of sperm nuclear proteins. Genome biology, 8(9), 1-8.] with UploadWizard