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Human taxonomy

(Redirected from Human subspecies)

Human taxonomy is the classification of the human species (systematic name Homo sapiens, Latin: "wise man") within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu (with some other research instead classifying idaltu and current humans as belonging to the same subspecies[1][2][3]).

Homo ("humans")
Temporal range: Piacenzian-Present, 2.865–0 Ma
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Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Infraorder: Simiiformes
Family: Hominidae
Subfamily: Homininae
Tribe: Hominini
Subtribe: Hominina
Genus: Homo
Linnaeus, 1758
Type species
Homo sapiens
Linnaeus, 1758
Species

other species or subspecies suggested

Synonyms
Synonyms
  • Africanthropus Dreyer, 1935
  • Atlanthropus Arambourg, 1954
  • Cyphanthropus Pycraft, 1928
  • Pithecanthropus Dubois, 1894
  • Protanthropus Haeckel, 1895
  • Sinanthropus Black, 1927
  • Tchadanthropus Coppens, 1965
  • Telanthropus Broom & Anderson 1949

Since the introduction of systematic names in the 18th century, knowledge of human evolution has increased drastically, and a number of intermediate taxa have been proposed in the 20th and early 21st centuries. The most widely accepted taxonomy grouping takes the genus Homo as originating between two and three million years ago, divided into at least two species, archaic Homo erectus and modern Homo sapiens, with about a dozen further suggestions for species without universal recognition.

The genus Homo is placed in the tribe Hominini alongside Pan (chimpanzees). The two genera are estimated to have diverged over an extended time of hybridization, spanning roughly 10 to 6 million years ago, with possible admixture as late as 4 million years ago. A subtribe of uncertain validity, grouping archaic "pre-human" or "para-human" species younger than the Homo-Pan split, is Australopithecina (proposed in 1939).

A proposal by Wood and Richmond (2000) would introduce Hominina as a subtribe alongside Australopithecina, with Homo the only known genus within Hominina. Alternatively, following Cela-Conde and Ayala (2003), the "pre-human" or "proto-human" genera of Australopithecus, Ardipithecus, Praeanthropus, and possibly Sahelanthropus, may be placed on equal footing alongside the genus Homo. An even more extreme view rejects the division of Pan and Homo as separate genera, which based on the Principle of Priority would imply the reclassification of chimpanzees as Homo paniscus (or similar).[4]

Categorizing humans based on phenotypes is a socially controversial subject. Biologists originally classified races as subspecies, but contemporary anthropologists reject the concept of race as a useful tool to understanding humanity, and instead view humanity as a complex, interrelated genetic continuum. Taxonomy of the hominins continues to evolve.[5][6]

History

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Further information: History of hominoid taxonomy, Hominini, Anthropomorpha, and Pithecanthropus
 
The taxonomic classification of humans following John Edward Gray (1825)

Human taxonomy on one hand involves the placement of humans within the taxonomy of the hominids (great apes), and on the other the division of archaic and modern humans into species and, if applicable, subspecies. Modern zoological taxonomy was developed by Carl Linnaeus during the 1730s to 1750s. He was the first to develop the idea that, like other biological entities, groups of people could too share taxonomic classifications.[7] He named the human species as Homo sapiens in 1758, as the only member species of the genus Homo, divided into several subspecies corresponding to the great races. The Latin noun homō (genitive hominis) means "human being". The systematic name Hominidae for the family of the great apes was introduced by John Edward Gray (1825).[8] Gray also supplied Hominini as the name of the tribe including both chimpanzees (genus Pan) and humans (genus Homo).

The discovery of the first extinct archaic human species from the fossil record dates to the mid 19th century: Homo neanderthalensis, classified in 1864. Since then, a number of other archaic species have been named, but there is no universal consensus as to their exact number. After the discovery of H. neanderthalensis, which even if "archaic" is recognizable as clearly human, late 19th to early 20th century anthropology for a time was occupied with finding the supposedly "missing link" between Homo and Pan. The "Piltdown Man" hoax of 1912 was the fraudulent presentation of such a transitional species. Since the mid-20th century, knowledge of the development of Hominini has become much more detailed, and taxonomical terminology has been altered a number of times to reflect this.

The introduction of Australopithecus as a third genus, alongside Homo and Pan, in the tribe Hominini is due to Raymond Dart (1925). Australopithecina as a subtribe containing Australopithecus as well as Paranthropus (Broom 1938) is a proposal by Gregory & Hellman (1939). More recently proposed additions to the Australopithecina subtribe include Ardipithecus (1995) and Kenyanthropus (2001). The position of Sahelanthropus (2002) relative to Australopithecina within Hominini is unclear. Cela-Conde and Ayala (2003) propose the recognition of Australopithecus, Ardipithecus, Praeanthropus, and Sahelanthropus (the latter incertae sedis)as separate genera.[9]

Other proposed genera, now mostly considered part of Homo, include: Pithecanthropus (Dubois, 1894), Protanthropus (Haeckel, 1895), Sinanthropus (Black, 1927), Cyphanthropus (Pycraft, 1928) Africanthropus (Dreyer, 1935),[10] Telanthropus (Broom & Anderson 1949), Atlanthropus (Arambourg, 1954), Tchadanthropus (Coppens, 1965).

The genus Homo has been taken to originate some two million years ago, since the discovery of stone tools in Olduvai Gorge, Tanzania, in the 1960s. Homo habilis (Leakey et al., 1964) would be the first "human" species (member of genus Homo) by definition, its type specimen being the OH 7 fossils. However, the discovery of more fossils of this type has opened up the debate on the delineation of H. habilis from Australopithecus. Especially, the LD 350-1 jawbone fossil discovered in 2013, dated to 2.8 Mya, has been argued as being transitional between the two.[11] It is also disputed whether H. habilis was the first hominin to use stone tools, as Australopithecus garhi, dated to c. 2.5 Mya, has been found along with stone tool implements.[12] Fossil KNM-ER 1470 (discovered in 1972, designated Pithecanthropus rudolfensis by Alekseyev 1978) is now seen as either a third early species of Homo (alongside H. habilis and H. erectus) at about 2 million years ago, or alternatively as transitional between Australopithecus and Homo.[13]

Wood and Richmond (2000) proposed that Gray's tribe Hominini ("hominins") be designated as comprising all species after the chimpanzee–human last common ancestor by definition, to the inclusion of Australopithecines and other possible pre-human or para-human species (such as Ardipithecus and Sahelanthropus) not known in Gray's time.[14] In this suggestion, the new subtribe of Hominina was to be designated as including the genus Homo exclusively, so that Hominini would have two subtribes, Australopithecina and Hominina, with the only known genus in Hominina being Homo. Orrorin (2001) has been proposed as a possible ancestor of Hominina but not Australopithecina.[15]

Designations alternative to Hominina have been proposed: Australopithecinae (Gregory & Hellman 1939) and Preanthropinae (Cela-Conde & Altaba 2002);[16][17][18]

Species

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Main article: Homo

At least a dozen species of Homo other than Homo sapiens have been proposed, with varying degrees of consensus. Homo erectus is widely recognized as the species directly ancestral to Homo sapiens.[citation needed] Most other proposed species are proposed as alternatively belonging to either Homo erectus or Homo sapiens as a subspecies. This concerns Homo ergaster in particular.[19][20] One proposal divides Homo erectus into an African and an Asian variety; the African is Homo ergaster, and the Asian is Homo erectus sensu stricto. (Inclusion of Homo ergaster with Asian Homo erectus is Homo erectus sensu lato.)[21] There appears to be a recent trend, with the availability of ever more difficult-to-classify fossils such as the Dmanisi skulls (2013) or Homo naledi fossils (2015) to subsume all archaic varieties under Homo erectus.[22][23][24]

Comparative table of Homo lineages
Lineages Temporal range
(kya) 		Habitat Adult height Adult mass Cranial capacity
(cm3) 		Fossil record Discovery Publication
of name
H. habilis
membership in Homo uncertain 		2,100–1,500[a][b] Tanzania 110–140 cm (3 ft 7 in – 4 ft 7 in) 33–55 kg (73–121 lb) 510–660 Many 1960 1964
H. rudolfensis
membership in Homo uncertain 		1,900 Kenya 700 2 sites 1972 1986
H. gautengensis
also classified as H. habilis 		1,900–600 South Africa 100 cm (3 ft 3 in) 3 individuals[27][c] 2010 2010
H. erectus 1,900–140[28][d][29][e] Africa, Eurasia 180 cm (5 ft 11 in) 60 kg (130 lb) 850 (early) – 1,100 (late) Many[f][g] 1891 1892
H. ergaster
African H. erectus 		1,800–1,300[31] East and Southern Africa 700–850 Many 1949 1975
H. antecessor 1,200–800 Western Europe 175 cm (5 ft 9 in) 90 kg (200 lb) 1,000 2 sites 1994 1997
H. heidelbergensis
early H. neanderthalensis 		600–300[h] Europe, Africa 180 cm (5 ft 11 in) 90 kg (200 lb) 1,100–1,400 Many 1907 1908
H. cepranensis
a single fossil, possibly H. heidelbergensis 		c. 450[32] Italy 1,000 1 skull cap 1994 2003
H. longi 309–138[33] Northeast China 1,420[34] 1 individual 1933 2021
H. rhodesiensis
early H. sapiens 		c. 300 Zambia 1,300 Single or very few 1921 1921
H. naledi c. 300[35] South Africa 150 cm (4 ft 11 in) 45 kg (99 lb) 450 15 individuals 2013 2015
H. sapiens
(anatomically modern humans) 		c. 300–present[i] Worldwide 150–190 cm (4 ft 11 in – 6 ft 3 in) 50–100 kg (110–220 lb) 950–1,800 (extant) —— 1758
H. neanderthalensis
 240–40[38][j] Europe, Western Asia 170 cm (5 ft 7 in) 55–70 kg (121–154 lb)
(heavily built) 		1,200–1,900 Many 1829 1864
H. floresiensis
classification uncertain 		190–50 Indonesia 100 cm (3 ft 3 in) 25 kg (55 lb) 400 7 individuals 2003 2004
Nesher Ramla Homo
classification uncertain 		140–120 Israel several individuals 2021
H. tsaichangensis
possibly H. erectus or Denisova 		c. 100[k] Taiwan 1 individual 2008(?) 2015
H. luzonensis
 c. 67[41][42] Philippines 3 individuals 2007 2019
Denisova hominin 40 Siberia 2 sites 2000
 2010[l]

Subspecies

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Homo sapiens subspecies

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Further information: Interbreeding between archaic and modern humans, Early modern human, and Race and genetics
 
1737 painting of Carl Linnaeus wearing a traditional Sami costume. Linnaeus is sometimes named as the lectotype of both H. sapiens and H. s. sapiens.[43]

The recognition or nonrecognition of subspecies of Homo sapiens has a complicated history. The rank of subspecies in zoology is introduced for convenience, and not by objective criteria, based on pragmatic consideration of factors such as geographic isolation and sexual selection. The informal taxonomic rank of race is variously considered equivalent or subordinate to the rank of subspecies, and the division of anatomically modern humans (H. sapiens) into subspecies is closely tied to the recognition of major racial groupings based on human genetic variation.

A subspecies cannot be recognized independently: a species will either be recognized as having no subspecies at all or at least two (including any that are extinct). Therefore, the designation of an extant subspecies Homo sapiens sapiens only makes sense if at least one other subspecies is recognized. H. s. sapiens is attributed to "Linnaeus (1758)" by the taxonomic Principle of Coordination.[44] During the 19th to mid-20th century, it was common practice to classify the major divisions of extant H. sapiens as subspecies, following Linnaeus (1758), who had recognized H. s. americanus, H. s. europaeus, H. s. asiaticus and H. s. afer as grouping the native populations of the Americas, West Eurasia, East Asia and Sub-Saharan Africa, respectively. Linnaeus also included H. s. ferus, for the "wild" form which he identified with feral children, and two other "wild" forms for reported specimens now considered very dubious (see cryptozoology), H. s. monstrosus and H. s. troglodytes.[45]

There were variations and additions to the categories of Linnaeus, such as H. s. tasmanianus for the native population of Australia.[46] Bory de St. Vincent in his Essai sur l'Homme (1825) extended Linnaeus's "racial" categories to as many as fifteen: Leiotrichi ("smooth-haired"): japeticus (with subraces), arabicus, iranicus, indicus, sinicus, hyperboreus, neptunianus, australasicus, columbicus, americanus, patagonicus; Oulotrichi ("crisp-haired"): aethiopicus, cafer, hottentotus, melaninus.[47] Similarly, Georges Vacher de Lapouge (1899) also had categories based on race, such as priscus, spelaeus (etc.).

Homo sapiens neanderthalensis was proposed by King (1864) as an alternative to Homo neanderthalensis.[48] There have been "taxonomic wars" over whether Neanderthals were a separate species since their discovery in the 1860s. Pääbo (2014) frames this as a debate that is unresolvable in principle, "since there is no definition of species perfectly describing the case."[49] Louis Lartet (1869) proposed Homo sapiens fossilis based on the Cro-Magnon fossils.

There are a number of proposals of extinct varieties of Homo sapiens made in the 20th century. Many of the original proposals were not using explicit trinomial nomenclature, even though they are still cited as valid synonyms of H. sapiens by Wilson & Reeder (2005).[50] These include: Homo grimaldii (Lapouge, 1906), Homo aurignacensis hauseri (Klaatsch & Hauser, 1910), Notanthropus eurafricanus (Sergi, 1911), Homo fossilis infrasp. proto-aethiopicus (Giuffrida-Ruggeri, 1915), Telanthropus capensis (Broom, 1917),[51] Homo wadjakensis (Dubois, 1921), Homo sapiens cro-magnonensis, Homo sapiens grimaldiensis (Gregory, 1921), Homo drennani (Kleinschmidt, 1931),[52] Homo galilensis (Joleaud, 1931) = Paleanthropus palestinus (McCown & Keith, 1932).[53] Rightmire (1983) proposed Homo sapiens rhodesiensis.[54]

After World War II, the practice of dividing extant populations of Homo sapiens into subspecies declined. An early authority explicitly avoiding the division of H. sapiens into subspecies was Grzimeks Tierleben, published 1967–1972.[55] A late example of an academic authority proposing that the human racial groups should be considered taxonomical subspecies is John Baker (1974).[56] The trinomial nomenclature Homo sapiens sapiens became popular for "modern humans" in the context of Neanderthals being considered a subspecies of H. sapiens in the second half of the 20th century. Derived from the convention, widespread in the 1980s, of considering two subspecies, H. s. neanderthalensis and H. s. sapiens, the explicit claim that "H. s. sapiens is the only extant human subspecies" appears in the early 1990s.[57]

Since the 2000s, the extinct Homo sapiens idaltu (White et al., 2003) has gained wide recognition as a subspecies of Homo sapiens, but even in this case there is a dissenting view arguing that "the skulls may not be distinctive enough to warrant a new subspecies name".[58] H. s. neanderthalensis and H. s. rhodesiensis continue to be considered separate species by some authorities, but the 2010s discovery of genetic evidence of archaic human admixture with modern humans has reopened the details of taxonomy of archaic humans.[59]

Homo erectus subspecies

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Further information: Homo erectus § Descendants and subspecies

Homo erectus since its introduction in 1892 has been divided into numerous subspecies, many of them formerly considered individual species of Homo. None of these subspecies have universal consensus among paleontologists.

See also

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Footnotes

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  1. ^ Confirmed H. habilis fossils are dated to between 2.1 and 1.5 million years ago. This date range overlaps with the emergence of Homo erectus.[25][26]
  2. ^ Hominins with "proto-Homo" traits may have lived as early as 2.8 million years ago, as suggested by a fossil jawbone classified as transitional between Australopithecus and Homo discovered in 2015.
  3. ^ A species proposed in 2010 based on the fossil remains of three individuals dated between 1.9 and 0.6 million years ago. The same fossils were also classified as H. habilis, H. ergaster or Australopithecus by other anthropologists.
  4. ^ H. erectus may have appeared some 2 million years ago. Fossils dated to as much as 1.8 million years ago have been found both in Africa and in Southeast Asia, and the oldest fossils by a narrow margin (1.85 to 1.77 million years ago) were found in the Caucasus, so that it is unclear whether H. erectus emerged in Africa and migrated to Eurasia, or if, conversely, it evolved in Eurasia and migrated back to Africa.
  5. ^ Homo erectus soloensis, found in Java, is considered the latest known survival of H. erectus. Formerly dated to as late as 50,000 to 40,000 years ago, a 2011 study pushed back the date of its extinction of H. e. soloensis to 143,000 years ago at the latest, more likely before 550,000 years ago. [30]
  6. ^ Now also included in H. erectus are Peking Man (formerly Sinanthropus pekinensis) and Java Man (formerly Pithecanthropus erectus).
  7. ^ H. erectus is now grouped into various subspecies, including Homo erectus erectus, Homo erectus yuanmouensis, Homo erectus lantianensis, Homo erectus nankinensis, Homo erectus pekinensis, Homo erectus palaeojavanicus, Homo erectus soloensis, Homo erectus tautavelensis, Homo erectus georgicus. The distinction from descendant species such as Homo ergaster, Homo floresiensis, Homo antecessor, Homo heidelbergensis and indeed Homo sapiens is not entirely clear.
  8. ^ The type fossil is Mauer 1, dated to ca. 0.6 million years ago. The transition from H. heidelbergensis to H. neanderthalensis between 300 and 243 thousand years ago is conventional, and makes use of the fact that there is no known fossil in this period. Examples of H. heidelbergensis are fossils found at Bilzingsleben (also classified as Homo erectus bilzingslebensis).
  9. ^ The age of H. sapiens has long been assumed to be close to 200,000 years, but since 2017 there have been a number of suggestions extending this time to as high as 300,000 years. In 2017, fossils found in Jebel Irhoud (Morocco) suggest that Homo sapiens may have speciated by as early as 315,000 years ago.[36] Genetic evidence has been adduced for an age of roughly 270,000 years.[37]
  10. ^ The first humans with "proto-Neanderthal traits" lived in Eurasia as early as 0.6 to 0.35 million years ago (classified as H. heidelbergensis, also called a chronospecies because it represents a chronological grouping rather than being based on clear morphological distinctions from either H. erectus or H. neanderthalensis). There is a fossil gap in Europe between 300 and 243 kya, and by convention, fossils younger than 243 kya are called "Neanderthal".[39]
  11. ^ younger than 450 kya, either between 190–130 or between 70–10 kya[40]
  12. ^ provisional names Homo sp. Altai or Homo sapiens ssp. Denisova.

References

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  39. ^ Dean D, Hublin JJ, Holloway R, Ziegler R (May 1998). "On the phylogenetic position of the pre-Neandertal specimen from Reilingen, Germany". Journal of Human Evolution. 34 (5): 485–508. doi:10.1006/jhev.1998.0214. PMID 9614635.
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  41. ^ Détroit F, Mijares AS, Corny J, Daver G, Zanolli C, Dizon E, et al. (April 2019). "A new species of Homo from the Late Pleistocene of the Philippines" (PDF). Nature. 568 (7751): 181–186. Bibcode:2019Natur.568..181D. doi:10.1038/s41586-019-1067-9. PMID 30971845. S2CID 106411053.
  42. ^ Zimmer C (10 April 2019). "A new human species once lived in this Philippine cave – Archaeologists in Luzon Island have turned up the bones of a distantly related species, Homo luzonensis, further expanding the human family tree". The New York Times. Retrieved 10 April 2019.
  43. ^ Ralph, Bob (February 19, 1987). "Conforming to type". New Scientist. No. 1548. p. 59. as far as I know, there is no type material for Homo sapiens. To be fair to Linnaeus, the practice of setting type specimens aside doesn't seem to have developed until a century or so later.
  44. ^ "article 46.1". ICZN glossary (4th ed.). International Code of Zoological Nomenclature. Archived from the original on 2016-03-03. Retrieved 2018-06-04. Statement of the Principle of Coordination applied to species-group names. A name established for a taxon at either rank in the species group is deemed to have been simultaneously established by the same author for a taxon at the other rank in the group; both nominal taxa have the same name-bearing type, whether that type was fixed originally or subsequently. Homo sapiens sapiens is rarely used before the 1940s. In 1946, John Wendell Bailey attributes the name to Linnaeus (1758) explicitly: "Linnaeus. Syst. Nat. ed. 10, Vol. 1. pp. 20, 21, 22, lists five races of man, viz: Homo sapiens sapiens (white — Caucasian) [...]", This is a misattribution, but H. s. sapiens has since often been attributed to Linnaeus. In actual fact, Linnaeus, Syst. Nat. ed. 10 Vol. 1. p. 21 does not have Homo sapiens sapiens, the "white" or "Caucasian" race being instead called Homo sapiens Europaeus. This is explicitly pointed out in Bulletin der Schweizerische Gesellschaft für Anthropologie und Ethnologie Volume 21 (1944), p. 18 (arguing not against H. s. sapiens but against "H. s. albus L." proposed by von Eickstedt and Peters): "die europide Rassengruppe, als Subspecies aufgefasst, [würde] Homo sapiens eurpoaeus L. heissen" ("the Europid racial group, considered as a subspecies, would be named H. s. europeaeus L."). See also: John R. Baker, Race, Oxford University Press (1974), 205.
  45. ^ Linné, Carl von (1758). Systema naturæ. Regnum animale (10 ed.). pp. 18ff.
  46. ^ See e.g. John Wendell Bailey, The Mammals of Virginia (1946), p. 356.; Journal of Mammalogy 26-27 (1945), p. 359.; J. Desmond Clark (ed.), The Cambridge History of Africa, Cambridge University Press (1982), p. 141 (with references).
  47. ^ Annals of Philosophy 11, London (1826), p. 71
  48. ^ Frederick S. Szalay, Eric Delson, Evolutionary History of the Primates (2013), 508
  49. ^ Pääbo, Svante (2014). Neanderthal Man: In Search of Lost Genomes. New York: Basic Books. p. 237.
  50. ^ Groves, C. P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press. ISBN 0-801-88221-4. OCLC 62265494.
  51. ^ Harrison, T. (2013). Kimbel, William H.; Martin, Lawrence B. (eds.). Species, Species Concepts and Primate Evolution. Springer. p. 361. ISBN 9781489937452.
  52. ^ M. R. Drennan, "An Australoid Skull from the Cape Flats", The Journal of the Royal Anthropological Institute of Great Britain and Ireland Vol. 59 (Jul. - Dec., 1929), 417-427.
  53. ^ among other names suggested for fossils later subsumed under neanderthalensis, see: Delson, Eric; Tattersall, Ian; Couvering, John Van; Brooks, Alison S. (2004). Encyclopedia of Human Evolution and Prehistory (Second ed.). Routledge. ISBN 9781135582272.
  54. ^ Rightmire, GP (June 3, 1983). "The Lake Ndutu cranium and early Homo sapiens in Africa". American Journal of Physical Anthropology. 61 (2): 245–54. doi:10.1002/ajpa.1330610214. PMID 6410925.
  55. ^ English translation (1972–1975): Grzimek's Animal Life Encyclopedia, Volume 11, p. 55.
  56. ^ John R. Baker, Race, Oxford University Press (1974).
  57. ^ Kitahara, Michio (1991). The tragedy of evolution: the human animal confronts modern society. p. xi. We are the only surviving subspecies of Homo sapiens
  58. ^ Stringer, Chris (June 12, 2003). "Human evolution: Out of Ethiopia". Nature. 423 (6941): 692–3, 695. Bibcode:2003Natur.423..692S. doi:10.1038/423692a. PMID 12802315. S2CID 26693109.
  59. ^ Hublin, J. J. (2009). "The origin of Neandertals". Proceedings of the National Academy of Sciences. 106 (38): 16022–7. Bibcode:2009PNAS..10616022H. doi:10.1073/pnas.0904119106. JSTOR 40485013. PMC 2752594. PMID 19805257. Harvati, K.; Frost, S.R.; McNulty, K.P. (2004). "Neanderthal taxonomy reconsidered: implications of 3D primate models of intra- and interspecific differences". Proceedings of the National Academy of Sciences. 101 (5): 1147–52. Bibcode:2004PNAS..101.1147H. doi:10.1073/pnas.0308085100. PMC 337021. PMID 14745010. "Homo neanderthalensis King, 1864". Wiley-Blackwell Encyclopedia of Human Evolution. Chichester, West Sussex: Wiley-Blackwell. 2013. pp. 328–331.
  60. ^ a b In the 1970s a tendency developed to regard the Javanese variety of H. erectus as a subspecies, Homo erectus erectus, with the Chinese variety being referred to as Homo erectus pekinensis. See: Sartono, S. (12 May 2011). "Implications arising from Pithecanthropus VIII". In Tuttle, Russell H. (ed.). Paleoanthropology: Morphology and Paleoecology. Walter de Gruyter. p. 328. ISBN 9783110810691.
  61. ^ Emanuel Vlček: Der fossile Mensch von Bilzingsleben (= Bilzingsleben. Bd. 6 = Beiträge zur Ur- und Frühgeschichte Mitteleuropas 35). Beier & Beran, Langenweißbach 2002.
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