Shri (named after Palden Lhamo, a Buddhist deity) is a genus of small dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, with fossils reported from Barun Goyot Formation in Khulsan, Mongolia. The type and only species, Shri devi, was described in 2021 by paleontologist Alan H. Turner and colleagues.[1]
| Shri Temporal range: Late Cretaceous (Maastrichtian), ~
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| Skeletal diagram of the holotype and ZPAL MgD-I/97 | |
Scientific classification 
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| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Dinosauria |
| Clade: | Saurischia |
| Clade: | Theropoda |
| Family: | †Dromaeosauridae |
| Clade: | †Eudromaeosauria |
| Subfamily: | †Velociraptorinae |
| Genus: | †Shri Turner et al, 2021 |
| Species: | †S. devi
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| Binomial name | |
| †Shri devi Turner et al., 2021
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History
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The holotype specimen of Shri is IGM 100/980. This specimen was discovered on 5 July 1991 by Mark Norell. It was nicknamed "Ichabodcraniosaurus" by Norell, as mentioned by Novacek (1996),[2] after Ichabod Crane, a fictional character haunted by a headless ghost, because it lacked the skull. In 1999 it was provisionally considered a specimen of Velociraptor mongoliensis.[3][2] It consists of a partially articulated individual that preserves the right hindlimb, the left tibiotarsus, as well as the pelvis and a series of cervical, dorsal, sacral and caudal vertebrae.[1]
In 2023, Polish paleontologist Łukasz Czepiński referred a new specimen to the genus, ZPAL MgD-I/97, represented by a partial skull and left hindlimb.[4] It was recovered from the Khulsan locality of the Barun Goyot Formation in 1970 during the Polish-Mongolian Paleontological Expeditions and initially assigned to Velociraptor.[5] Further examinations by Czepiński concluded it to represent an additional specimen of Shri based on pes (foot) morphology.[4]
Description
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Shri was mostly similar to Velociraptor mongoliensis in having a weak fourth trochanter, this is however, also shared with all other dromaeosaurids, and deep anterior pedicular fossae in the cervical vertebrae. Another distinguishing trait of Shri is that its epipophyses in the last four cervicals are not raised but instead are represented by rugose circular scars.[1]
Classification
editShri devi was entered into a phylogenetic analysis to test its relationships within Dromaeosauridae. It was found to be the sister taxon of Velociraptor mongoliensis based on the presence of a distinct ambiens tubercle that is located proximally on the anterior face of the pubis, a well-developed anterior tuberosity located high on the ischium, as well as a rounded ischial ridge that runs lengthwise. A cladogram of the phylogenetic analysis performed by the describers is shown below:[1]
References
edit- ^ a b c d Turner, Alan H.; Montanari, Shaena; Norell, Mark A. (2021). "A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia" (PDF). American Museum Novitates (3965): 1–48. doi:10.1206/3965.1. ISSN 0003-0082. S2CID 231597229.
- ^ a b Novacek, Michael J. (1996). Dinosaurs of the Flaming Cliffs. New York: Anchor Books. ISBN 0-385-47774-0.
- ^ Norell, Mark A.; Makovicky, Peter J. (1999). "Important features of the dromaeosaurid skeleton II: information from newly collected specimens of Velociraptor mongoliensis". American Museum Novitates (3282): 1–45. hdl:2246/3025.
- ^ a b Czepiński, Ł. (2023). "Skull of a dromaeosaurid dinosaur Shri devi from the Upper Cretaceous of the Gobi Desert suggests convergence to the North American forms" (PDF). Acta Palaeontologica Polonica. 68. doi:10.4202/app.01065.2023.
- ^ Barsbold, R.; Osmólska, H. (1999). "The skull of Velociraptor (Theropoda) from the Late Cretaceous of Mongolia" (PDF). Acta Palaeontologica Polonica. 44 (2): 189–219.
