The Cetaceans Portal
Cetacea (/sɪˈteɪʃə/; from Latin cetus 'whale', from Ancient Greek κῆτος (kêtos) 'huge fish, sea monster') is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver.
While the majority of cetaceans live in marine environments, a small number reside solely in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons.
Cetaceans are famous for their high intelligence, complex social behaviour, and the enormous size of some of the group's members. For example, the blue whale reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short tons), making it the largest animal ever known to have existed.
There are approximately 89 living species split into two parvorders: Odontoceti or toothed whales (containing porpoises, dolphins, other predatory whales like the beluga and the sperm whale, and the poorly understood beaked whales) and the filter feeding Mysticeti or baleen whales (which includes species like the blue whale, the humpback whale and the bowhead whale). Despite their highly modified bodies and carnivorous lifestyle, genetic and fossil evidence places cetaceans as nested within even-toed ungulates, most closely related to hippopotamus within the clade Whippomorpha. (Full article...)
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Pilot whales are cetaceans belonging to the genus Globicephala. The two extant species are the long-finned pilot whale (G. melas) and the short-finned pilot whale (G. macrorhynchus). The two are not readily distinguishable at sea, and analysis of the skulls is the best way to distinguish between the species. Between the two species, they range nearly worldwide, with long-finned pilot whales living in colder waters and short-finned pilot whales living in tropical and subtropical waters. Pilot whales are among the largest of the oceanic dolphins, exceeded in size only by the orca. They and other large members of the dolphin family are also known as blackfish.
Pilot whales feed primarily on squid, but will also hunt large demersal fish such as cod and turbot. They are highly social and may remain with their birth pod throughout their lifetime. Short-finned pilot whales are one of the few non-primate mammal species in which females go through menopause, and postreproductive females continue to contribute to their pod. Pilot whales are notorious for stranding themselves on beaches, but the reason behind this is not fully understood. Marine biologists have shed some light on the matter, suggesting that it is due to the mammals inner ear (their principal navigational sonar) being damaged from noise pollution in the ocean, such as from cargo ships or military exercises. The conservation status of short-finned and long-finned pilot whales has been determined to be least concern. (Full article...) -
The sei whale (/seɪ/ SAY, Norwegian: [sæɪ]; Balaenoptera borealis) is a baleen whale. It is one of ten rorqual species, and the third-largest member after the blue and fin whales. It can grow to 19.5 m (64 ft) in length and weigh as much as 28 t (28 long tons; 31 short tons). Two subspecies are recognized: B. b. borealis and B. b. schlegelii. The whale's ventral surface has sporadic markings ranging from light grey to white, and its body is usually dark steel grey in colour. It is among the fastest of all cetaceans, and can reach speeds of up to 50 km/h (31 mph) over short distances.
It inhabits most oceans and adjoining seas, and prefers deep offshore waters. It avoids polar and tropical waters and semi-enclosed bodies of water. The sei whale migrates annually from cool, subpolar waters in summer to temperate, subtropical waters in winter with a lifespan of 70 years. It is a filter feeder, with its diet consisting primarily of copepods, krill, and other zooplankton. It is typically solitary or can be found in groups numbering half a dozen. During the breeding period, a mating pair will remain together. Sei whale vocalizations usually lasts half a second, and occurs at 240–625 hertz. (Full article...) -
The anatomist John Struthers (at left, in top hat) with the Tay Whale at John Woods's yard, Dundee, 1884, photographed by George Washington Wilson
The Tay Whale, known locally as the Monster, was a humpback whale that swam into the Firth of Tay of eastern Scotland in 1883. It was harpooned in a hunt, but escaped, and was found floating dead off Stonehaven a week later. It was towed into Dundee by a showman, John Woods, and exhibited on a train tour of Scotland and England.
The Regius Professor of Anatomy at Aberdeen University, John Struthers dissected the whale, much of the time in public with a military band playing in the background, organised by Woods. The decomposing whale made Woods a great deal of money, and Struthers famous. (Full article...) -
The humpback whale (Megaptera novaeangliae) is a species of baleen whale. It is a rorqual (a member of the family Balaenopteridae) and is the only species in the genus Megaptera. Adults range in length from 14–17 m (46–56 ft) and weigh up to 40 metric tons (44 short tons). The humpback has a distinctive body shape, with long pectoral fins and tubercles on its head. It is known for breaching and other distinctive surface behaviors, making it popular with whale watchers. Males produce a complex song typically lasting 4 to 33 minutes.
Found in oceans and seas around the world, humpback whales typically migrate up to 16,000 km (9,900 mi) each year. They feed in polar waters and migrate to tropical or subtropical waters to breed and give birth. Their diet consists mostly of krill and small fish, and they use bubbles to catch prey. They are promiscuous breeders, with both sexes having multiple partners. Orcas are the main natural predators of humpback whales. (Full article...) -
Archaeopteryx is one of the most famous transitional fossils and gives evidence for the evolution of birds from theropod dinosaurs.
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors.
In 1859, when Charles Darwin's On the Origin of Species was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be urged against my theory," but he explained it by relating it to the extreme imperfection of the geological record. He noted the limited collections available at the time but described the available information as showing patterns that followed from his theory of descent with modification through natural selection. Indeed, Archaeopteryx was discovered just two years later, in 1861, and represents a classic transitional form between earlier, non-avian dinosaurs and birds. Many more transitional fossils have been discovered since then, and there is now abundant evidence of how all classes of vertebrates are related, including many transitional fossils. Specific examples of class-level transitions are: tetrapods and fish, birds and dinosaurs, and mammals and "mammal-like reptiles". (Full article...) -
Brygmophyseter, known as the biting sperm whale, is an extinct genus of toothed whale in the sperm whale family with one species, B. shigensis. When it was first described in 1994, the species was placed in the genus Scaldicetus based on tooth morphology, but this was later revised in 1995. In 2006, it was classified into the genus Naganocetus, which is considered to be a junior synonym. The only known specimen, a nearly complete skeleton, was dated to be around 16–15 million years old (middle Miocene). Brygmophyseter is thought to have been 6.5–7 meters (21–23 ft) long, and it probably had 11 or 12 teeth in the upper and lower jaws. Brygmophyseter is part of a group of macroraptorial sperm whales (often shortened to "raptorial") which tended to be apex predators using their large teeth to catch struggling prey such as whales. It had a spermaceti organ which was probably used for biosonar like in the modern sperm whale. The whale has made an appearance on The History Channel's TV series Jurassic Fight Club. (Full article...) -
Rice's whale (Balaenoptera ricei), also known as the Gulf of Mexico whale, is a species of baleen whale endemic to the northern Gulf of Mexico. Initially identified as a subpopulation of the Bryde's whale, genetic and skeletal studies found it to be a distinct species by 2021. In outward appearance, it is virtually identical to the Bryde's whale. Its body is streamlined and sleek, with a uniformly dark charcoal gray dorsal and pale to pinkish underside. A diagnostic feature often used by field scientists to distinguish Rice's whales from whales other than the Bryde's whale is the three prominent ridges that line the top of its head. The species can be distinguished from the Bryde's whale by the shape of the nasal bones, which have wider gaps due to a unique wrapping by the frontal bones, its unique vocal repertoire, and genetic differences.
It is a medium-sized baleen whale that grows up to 12.65 meters (41.5 ft) in length and weighs up to 13.87–27.2 metric tons (13.65–26.77 long tons; 15.29–29.98 short tons). The Rice's whale inhabits a restricted stretch along the continental slope in the northeastern part of the Gulf of Mexico between depths of 150–410 meters (490–1,350 ft) off the coast of western Florida, although some whales have been sighted in the northwestern portions and the species may have inhabited a wider distribution throughout the Gulf in historical times. It does not migrate but remains within this area year-round. Little is known about the feeding behavior of Rice's whales, but data from a tagged individual revealed a diel vertical diving pattern, in which the whale spends most of the day feeding at or near the seafloor at depths of up to 271 meters (889 ft) and night at the surface. The whale's diet remains unknown, but lanternfish and hatchetfish are suspected prey. (Full article...) -
The bottlenose dolphin is a toothed whale in the genus Tursiops. They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus contains three species: the common bottlenose dolphin (Tursiops truncatus), the Indo-Pacific bottlenose dolphin (Tursiops aduncus), and Tamanend's bottlenose dolphin (Tursiops erebennus). Others, like the Burrunan dolphin (Tursiops (aduncus) australis), may be alternately considered their own species or be subspecies of T. aduncus. Bottlenose dolphins inhabit warm and temperate seas worldwide, being found everywhere except for the Arctic and Antarctic Circle regions. Their name derives from the Latin tursio (dolphin) and truncatus for the truncated teeth (the type specimen was old and had worn down teeth; this is not a typical characteristic of most members of the species).
Numerous investigations of bottlenose dolphin intelligence have been conducted, examining mimicry, use of artificial language, object categorization, and self-recognition. They can use tools (sponging; using marine sponges to forage for food sources they normally could not access) and transmit cultural knowledge from generation to generation, and their considerable intelligence has driven interaction with humans. Bottlenose dolphins gained popularity from aquarium shows and television programs such as Flipper. They have also been trained by militaries to locate sea mines or detect and mark enemy divers. In some areas, they cooperate with local fishermen by driving fish into their nets and eating the fish that escape. Some encounters with humans are harmful to the dolphins: people hunt them for food, and dolphins are killed inadvertently as a bycatch of tuna fishing and by getting caught in crab traps. (Full article...) -
Porpoises (/ˈpɔːrpəsɪz/) are small dolphin-like cetaceans classified under the family Phocoenidae. Although similar in appearance to dolphins, they are more closely related to narwhals and belugas than to the true dolphins. There are eight extant species of porpoise, all among the smallest of the toothed whales. Porpoises are distinguished from dolphins by their flattened, spade-shaped teeth distinct from the conical teeth of dolphins, and lack of a pronounced beak, although some dolphins (e.g. Hector's dolphin) also lack a pronounced beak. Porpoises, and other cetaceans, belong to the clade Cetartiodactyla with even-toed ungulates.
Porpoises range in size from the vaquita, at 1.4 metres (4 feet 7 inches) in length and 54 kilograms (119 pounds) in weight, to the Dall's porpoise, at 2.3 m (7 ft 7 in) and 220 kg (490 lb). Several species exhibit sexual dimorphism in that the females are larger than males. They have streamlined bodies and two limbs that are modified into flippers. Porpoises use echolocation as their primary sensory system. Some species are well adapted for diving to great depths. As all cetaceans, they have a layer of fat, or blubber, under the skin to keep them warm in cold water. (Full article...) -
Granny (born c. 1936-1951 approx, died c. 2016), also known as J2, was a female orca of the J pod of southern resident orcas notable for her long life. Early estimates placed her birth in 1911, putting her at 105 years old at the time of her death. However, this estimate was later theorized to have been based on mistaken information and more recent studies put her at 65–80 years old. If she was 105, she would have been the oldest known orca at the time of her death. Granny lived in the northeast Pacific Ocean and coastal bays of Washington state and British Columbia. She was last seen on October 12, 2016, and was considered deceased by The Center for Whale Research in January 2017. (Full article...) -
Species of the infraorder Cetacea
The evolution of cetaceans is thought to have begun in the Indian subcontinent from even-toed ungulates (Artiodactyla) 50 million years ago (mya) and to have proceeded over a period of at least 15 million years. Cetaceans are fully aquatic mammals belonging to the order Artiodactyla and branched off from other artiodactyls around 50 mya. Cetaceans are thought to have evolved during the Eocene (56-34 mya), the second epoch of the present-extending Cenozoic Era. Molecular and morphological analyses suggest Cetacea share a relatively recent closest common ancestor with hippopotami and that they are sister groups. Being mammals, they surface to breathe air; they have five finger bones (even-toed) in their fins; they nurse their young; and, despite their fully aquatic life style, they retain many skeletal features from their terrestrial ancestors. Research conducted in the late 1970s in Pakistan revealed several stages in the transition of cetaceans from land to sea.
The two modern parvorders of cetaceans – Mysticeti (baleen whales) and Odontoceti (toothed whales) – are thought to have separated from each other around 28–33 mya in a second cetacean radiation, the first occurring with the archaeocetes. The adaptation of animal echolocation in toothed whales distinguishes them from fully aquatic archaeocetes and early baleen whales. The presence of baleen in baleen whales occurred gradually, with earlier varieties having very little baleen, and their size is linked to baleen dependence (and subsequent increase in filter feeding). (Full article...) -
The narwhal (Monodon monoceros) is a species of toothed whale native to the Arctic. It is the only member of the genus Monodon and one of two living representatives of the family Monodontidae. The narwhal has a similar build to the closely related beluga whale, with which it overlaps in range and can interbreed. It is a stocky cetacean with a relatively blunt snout, a large melon, and a shallow ridge in place of a dorsal fin. Males of this species have a large 1.5–3.0 m (4 ft 11 in – 9 ft 10 in) long tusk, which is a left protruded canine thought to serve as a weapon or as a tool for feeding, attracting mates or sensing water salinity. Specially adapted slow-twitch muscles, along with the jointed neck vertebrae and shallow dorsal ridge allow for easy movement through the Arctic environment, where the narwhal spends extended periods at great depths.
The narwhal inhabits the Arctic waters of Canada, Greenland and Russia. Every year, it migrates to ice-free summering grounds, usually in shallow waters, and often returns to the same sites in subsequent years. Its diet mainly consists of polar and Arctic cod, Greenland halibut, cuttlefish, shrimp, and armhook squid. Plunging at depths of up to 2,370 m (7,780 ft), the narwhal is among the deepest-diving cetaceans. It travels in groups of three to eight, with aggregations of up to 1,000 occurring in the summer months. It mates in the offshore pack ice from March to May, and the young are born in July or August of the following year. When communicating, a variety of clicks, whistles and knocks are used. (Full article...) -
River dolphins are a polyphyletic group of fully aquatic mammals that reside exclusively in freshwater or brackish water. They are an informal grouping of dolphins, which itself is a paraphyletic group within the infraorder Cetacea. Extant river dolphins are placed in two superfamilies, Platanistoidea and Inioidea. They comprise the families Platanistidae (the South Asian dolphins), the possibly extinct Lipotidae (Yangtze River dolphin), Iniidae (the Amazonian dolphins) and Pontoporiidae. There are five extant species of river dolphins. River dolphins, alongside other cetaceans, belong to the clade Artiodactyla, with even-toed ungulates, and their closest living relatives the hippopotamuses, from which they diverged about 40 million years ago. Specific types of dolphins can be pink.
River dolphins are relatively small compared to other dolphins, having evolved to survive in warm, shallow water and strong river currents. They range in size from the 5-foot (1.5 m) long South Asian river dolphin to the 8-foot (2.4 m) and 220-pound (100 kg) Amazon river dolphin. Several species exhibit sexual dimorphism, in that the females are larger than the males. They have streamlined bodies and two limbs that are modified into flippers. River dolphins use their conical-shaped teeth and long beaks to capture fast-moving prey in murky water. They have well-developed hearing that is adapted for both air and water; they do not really rely on vision since the water they swim in is usually very muddy. Instead, they tend to rely on echolocation when hunting and navigating. These species are well-adapted to living in warm, shallow waters, and, unlike other cetaceans, have little to no blubber. (Full article...) -
Whaling in the Faroe Islands, or grindadráp (from the Faroese terms grindhvalur, meaning pilot whale, and dráp, meaning killing), is a type of drive hunting that involves herding various species of whales and dolphins, but primarily pilot whales, into shallow bays to be beached, killed, and butchered. Each year, an average of around 700 long-finned pilot whales and several hundred Atlantic white-sided dolphins are caught over the course of the hunt season during the summer.
The practice dates back to the 9th century, and many Faroe Islanders consider eating whales to be an important part of their history. Since 1948, the hunt has been regulated by the Faroese authorities, required its participants to be trained, involved modern boats and communications, and been supervised by police. (Full article...) -
Orcinus citoniensis is an extinct species of orca identified in the Late Pliocene of Italy and the Early Pleistocene of England. It was smaller than the modern killer whale (O. orca), 4 m (13 ft) versus 7 to 10 m (23 to 33 ft), and had around 8 more teeth in its jaw. It may have resembled the modern killer whale in appearance, and could represent a transitional species between the modern killer whale and other dolphins. O. citoniensis could have hunted fish and squid in pods, and coexisted with other large predators of the time such as the orcinine Hemisyntrachelus and the extinct shark Otodus megalodon. (Full article...)
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Photo credit: Cornelia Oedekoven
Many whales are known to breach the waters surface. The act of leaping generates more power than any other act performed by a non-human animal.
Some whales, such as Sperm Whales, perform a breach by travelling vertically upwards from depth, and heading straight out of the water. Others, such as the Humpback Whale (pictured), travel close to the surface and parallel to it, and then jerk upwards at full speed to perform a breach. In a typical breach, as performed by a Humpback or Right Whale, the whale clears the water at an angle of about 30° to the horizontal.
More on whale behaviour
More did you know...
- ...cetaceans have evolved from land mammals. Evidence of this is seen in the vestigial hip bones, as well in the pentadactyl ("five-fingered") dorsal flippers/fins.
- ...Herman Melville included an objective study of the properties of whales in Moby-Dick.
- ...a male dolphin named Cinderella was unofficially married to a human woman.
- ...sound made by whales can be extremely loud, with 163 decibels recorded.
- ...about 2000 whales may be beached each year.
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General images -
The following are images from various cetacean-related articles on Wikipedia.
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Common minke whale illustration -
South Atlantic right whale illustration -
Species of the infraorder Cetacea (from Evolution of cetaceans) -
Pygmy right whale illustration -
Restoration of Janjucetus hunderi (from Baleen whale) -
Japanese scientific whaling on a pair of Antarctic minke whales (from Baleen whale) -
Sei whale illustration -
Anatomy of the bottlenose dolphin (from Toothed whale) -
Archaeocetes (like this Basilosaurus) had a heterodont dentition (from Evolution of cetaceans) -
Biosonar by cetaceans (from Toothed whale) -
Fin whale illustration -
Blue whale illustration -
Possible relationships between cetaceans and other ungulate groups. (from Evolution of cetaceans) -
The remains of a North Atlantic right whale after it collided with a ship propeller -
North Atlantic right whale illustration -
Female right whale with calf -
The heart of a blue whale with a person standing next to it -
Features of a sperm whale skeleton (from Toothed whale) -
Bryde's whale (rorqual) (from Baleen whale) -
Gray whale illustration -
Orca hunting a Weddell seal (from Toothed whale) -
Their eyes are relatively small for their size. -
A humpback whale skeleton. Notice how the jaw is split into two. -
A phylogenetic tree showing the relationships among cetacean families. (from Evolution of cetaceans) -
A protest against Japan's scientific whaling -
Megalodon hunted Miocene baleen whales (from Baleen whale) -
Humpback whale illustration -
Diagram illustrating sound generation, propagation and reception in a toothed whale. Outgoing sounds are red and incoming ones are green (from Toothed whale) -
Antarctic minke whale illustration -
Orange whale lice on a right whale -
Pakicetus attocki skeleton (from Evolution of cetaceans) -
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Atlantic white-sided dolphin caught in a drive hunt in Hvalba on the Faroe Islands being taken away with a forklift (from Toothed whale) -
Acrophyseter skull (from Evolution of cetaceans) -
An orca by the name of Ulises performing at SeaWorld, 2009 (from Toothed whale) -
World population graph of blue whales -
The skeleton of a bowhead whale with the hind limb and pelvic bone structure circled in red. This bone structure stays internal during the entire life of the species. (from Evolution of cetaceans) -
Humpback whales lunge-feeding in the course of bubble net fishing -
Baleen whales vary considerably in size and shape, depending on their feeding behavior. (from Baleen whale) -
Eden's whale illustration -
Fossil of Squalodon (from Toothed whale) -
Dolphins (aquatic mammals) and ichthyosaurs (extinct marine reptiles) share a number of unique adaptations for fully aquatic lifestyle and are frequently used as extreme examples of convergent evolution (from Evolution of cetaceans) -
Archaeomysticetes, like Janjucetus, had teeth. -
Bowhead whale illustration -
Accessory baleen plates taper off into small hairs (from Baleen whale) -
The nose of the whale is filled with a waxy substance that was widely used in candles, oil lamps, and lubricants (from Toothed whale) -
Spectrogram of dolphin vocalizations. Whistles, whines, and clicks are visible as upside down V's, horizontal striations, and vertical lines, respectively. (from Toothed whale)
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Did you know (auto-generated)
- ... that Celia Kaye won the Golden Globe Award for Most Promising Newcomer in 1965 for her starring role in Island of the Blue Dolphins?
- ... that the South Asian river dolphin is nearly blind and relies on echolocation for navigation?
- ... that one of the first researchers to propose dolphin-assisted therapy for humans later renounced it?
- ... that one can swim with humpback whales in the Niue Nukutuluea Multiple-Use Marine Park?
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