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The Cenozoic Portal

Introduction

The Cenozoic (/ˌsiːnəˈzoʊ.ɪk, ˌsɛn-/ SEE-nə-ZOH-ik, SEN-ə-; lit. 'new life') is Earth's current geological era, representing the last 66 million years of Earth's history. It is characterized by the dominance of insects, mammals, birds and angiosperms (flowering plants). It is the latest of three geological eras of the Phanerozoic Eon, preceded by the Mesozoic and Paleozoic. The Cenozoic started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor.

The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammals – the eutherians (placentals) in the Northern Hemisphere and the metatherians (marsupials, now mainly restricted to Australia and to some extent South America) in the Southern Hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that large mammals and birds dominated life on Earth. The continents also moved into their current positions during this era.

The climate during the early Cenozoic was warmer than today, particularly during the Paleocene–Eocene Thermal Maximum. However, the Eocene to Oligocene transition and the Quaternary glaciation dried and cooled Earth. (Full article...)

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Selected article on the Cenozoic world and its legacies

Archaeoindris fontoynontii skull

Archaeoindris fontoynontii is an extinct giant lemur and the largest primate known to have evolved on Madagascar, comparable in size to a male gorilla. It belonged to a family of extinct lemurs known as "sloth lemurs" (Palaeopropithecidae) and, because of its extremely large size, it has been compared to the ground sloths that once roamed North and South America. It was most closely related to Palaeopropithecus, the second largest type of sloth lemur. Along with the other sloth lemurs, Archaeoindris was related to the living indri, sifakas, and woolly lemurs, as well as the recently extinct monkey lemurs (Archaeolemuridae). The genus, Archaeoindris, translates to "ancient indri-like lemur", even though it probably became extinct recently, around 350 BCE.

Archaeoindris was first described by Herbert F. Standing in 1909 on the basis of subfossil fragmentary jaws, although Charles Lamberton later discovered a complete skull. Only six bones from the lower skeleton have been found and excavations in the 1980s offered no leads for new finds. Its remains have been found at only one location: Ampasambazimba, a subfossil site in central Madagascar. Following its initial discovery, some subfossil remains of Megaladapis grandidieri (a type of extinct koala lemur) were mistakenly associated with Archaeoindris, while smaller leg bones from a juvenile and a massive adult leg bone were erroneously assumed to belong to two separate species. These errors were gradually corrected between the 1930s and 1980s.

The skeleton of Archaeoindris was massive and robust, and shared many traits with that of Palaeopropithecus. The arms were longer than the legs, but no hand or foot bones have been found for comparison with the other sloth lemurs. Size estimates based on the limited remains have varied widely, ranging as high as 244 kilograms (538 pounds), but the most thorough statistical investigation using regression analyses predicts a mass of 160 kg (350 lb).

Misattributions and limited remains have resulted in varying opinions about the way Archaeoindris moved in its environment, ranging from tree-dwelling to ground-dwelling. Its skeleton suggests it was a deliberate climber that visited the ground to travel.

The diet of Archaeoindris was mostly leaves, and its habitat—prior to human arrival—was a mix of woodlands, bushlands, and savanna, rich in lemur diversity. Today, the region is dominated by grasslands and lemur diversity is very low in the nearest protected area, Ambohitantely Special Reserve. Although it was a rare lemur, it was still extant when humans first arrived on Madagascar, and it would have been vulnerable to hunting and habitat loss. (Full article...)

Selected article on the Cenozoic in human science, culture and economics

Evolutionary thought, the recognition that species change over time and the perceived understanding of how such processes work, has roots in antiquity—in the ideas of the ancient Greeks, Romans, Chinese, Church Fathers as well as in medieval Islamic science. With the beginnings of modern biological taxonomy in the late 17th century, two opposed ideas influenced Western biological thinking: essentialism, the belief that every species has essential characteristics that are unalterable, a concept which had developed from medieval Aristotelian metaphysics, and that fit well with natural theology; and the development of the new anti-Aristotelian approach to modern science: as the Enlightenment progressed, evolutionary cosmology and the mechanical philosophy spread from the physical sciences to natural history. Naturalists began to focus on the variability of species; the emergence of palaeontology with the concept of extinction further undermined static views of nature. In the early 19th century prior to Darwinism, Jean-Baptiste Lamarck (1744–1829) proposed his theory of the transmutation of species, the first fully formed theory of evolution.

In 1858 Charles Darwin and Alfred Russel Wallace published a new evolutionary theory, explained in detail in Darwin's On the Origin of Species (1859). Darwin's theory, originally called descent with modification is known contemporarily as Darwinism or Darwinian theory. Unlike Lamarck, Darwin proposed common descent and a branching tree of life, meaning that two very different species could share a common ancestor. Darwin based his theory on the idea of natural selection: it synthesized a broad range of evidence from animal husbandry, biogeography, geology, morphology, and embryology. Debate over Darwin's work led to the rapid acceptance of the general concept of evolution, but the specific mechanism he proposed, natural selection, was not widely accepted until it was revived by developments in biology that occurred during the 1920s through the 1940s. Before that time most biologists regarded other factors as responsible for evolution. Alternatives to natural selection suggested during "the eclipse of Darwinism" (c. 1880 to 1920) included inheritance of acquired characteristics (neo-Lamarckism), an innate drive for change (orthogenesis), and sudden large mutations (saltationism). Mendelian genetics, a series of 19th-century experiments with pea plant variations rediscovered in 1900, was integrated with natural selection by Ronald Fisher, J. B. S. Haldane, and Sewall Wright during the 1910s to 1930s, and resulted in the founding of the new discipline of population genetics. During the 1930s and 1940s population genetics became integrated with other biological fields, resulting in a widely applicable theory of evolution that encompassed much of biology—the modern synthesis.

Following the establishment of evolutionary biology, studies of mutation and genetic diversity in natural populations, combined with biogeography and systematics, led to sophisticated mathematical and causal models of evolution. Palaeontology and comparative anatomy allowed more detailed reconstructions of the evolutionary history of life. After the rise of molecular genetics in the 1950s, the field of molecular evolution developed, based on protein sequences and immunological tests, and later incorporating RNA and DNA studies. The gene-centred view of evolution rose to prominence in the 1960s, followed by the neutral theory of molecular evolution, sparking debates over adaptationism, the unit of selection, and the relative importance of genetic drift versus natural selection as causes of evolution. In the late 20th-century, DNA sequencing led to molecular phylogenetics and the reorganization of the tree of life into the three-domain system by Carl Woese. In addition, the newly recognized factors of symbiogenesis and horizontal gene transfer introduced yet more complexity into evolutionary theory. Discoveries in evolutionary biology have made a significant impact not just within the traditional branches of biology, but also in other academic disciplines (for example: anthropology and psychology) and on society at large. (Full article...)

Selected images

Image 1Prolibytherium ("Before Libya's Beast") is an extinct climacoceratid artiodactyl ungulate native to Early Miocene North Africa and Pakistan
Image 2Adcrocuta is an extinct genus of terrestrial carnivore in the family Hyaenidae that lived during the Miocene period
Image 3Palaeotragus ("ancient goat") is a genus of very large, primitive, okapi-like giraffid from the Miocene of Africa, Eurasia.
Image 4Mimoperadectes is an extinct genus of didelphimorph marsupial from the Eocene of North America
Image 5Victoriapithecus macinnesi was a primate from middle Miocene central Africa
Image 6Morotopithecus bishopi is a species of fossil ape discovered in Miocene-age deposits of Moroto, Uganda.
Image 7Indarctos is a genus of mammals of the bear family, Ursidae, endemic to North America, Europe and Asia during the Miocene
Image 8Stegotetrabelodon is an extinct genus of primitive elephant with gomphothere-like anatomical features from the Late Miocene to Early Pliocene of Africa and Eurasia
Image 9Platybelodon ("flat-spear tusk") was a genus of large herbivorous mammals related to the elephant (order Proboscidea). It lived during the late Miocene Epoch in Asia and the Caucasus.
Image 10Deinotherium was a large prehistoric relative of modern-day elephants that appeared in the Middle Miocene and survived until the Early Pleistocene
Image 11Proconsul is an extinct genus of primates that existed from 23 to 25 million years ago during the Miocene epoch
Image 12Fossil tooth of Hyainailurus, an extinct genus of hyaenodont mammal of the family Hyainailouridae that lived during the Miocene