The Ashley Formation is a geologic formation in South Carolina. It preserves fossils dating back to the Paleogene period.
Ashley Formation | |
---|---|
Stratigraphic range: Lower Oligocene (Rupelian) | |
Type | Geological formation |
Sub-units | Gettysville Member, Runnymeade Marl Member and Givhans Ferry Member.[1] |
Location | |
Region | South Carolina |
Country | United States |
Vertebrate fauna
editMammals
editCarnivorans | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Phocidae | Gen. et. sp. indeterminate | Proximal portion of a right femur (ChM PV5713).[2] | "A taxon closely comparable to the most specialized phocid, the modern genus Cystophora". |
Cetaceans | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Agorophius | A. pygmaeus | Underwater cliff face from the base of the formation.[3] | A skull.[3] | ||
Albertocetus | A. meffordorum | Givhans Ferry Member and Runnymede Marl Member.[4] | A partial skeleton and an isolated braincase.[4] | A xenorophid. | |
Ankylorhiza | A. tiedemani | AMNH 10445 (partial rostrum) and CCNHM 220 (partial skull and associated vertebrae).[5] | An odontocete. | ||
Ashleycetus | A. planicapitis | Near Charleston, South Carolina.[6] | A partial skull.[6] | ||
Coronodon | C. havensteini | "Wando River near Highway 41 Bridge, Berkeley County, South Carolina".[7] | A basal mysticete. | ||
Ediscetus | E. osbornei | "Slightly outside the odontocete crown group".[8] | |||
Micromysticetus | M. rothauseni | 2 braincases without rostra.[9] | |||
Saurocetus | S. gibbesi | [10] | A cheek tooth (MCZ 8760).[11] | ||
Xenorophus | X. sloanii | Incomplete skull.[12] | A xenorophid. |
Sirenians | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Crenatosiren | C. olseni | [13] | A dugongid also found in the Chandler Bridge and Parachucla formations. | ||
Stegosiren | S. macei | [14] | A dugongid also found in the Chandler Bridge Formation. |
Reptiles
editColor key
|
Notes Uncertain or tentative taxa are in small text; |
Crocodilians | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Gavialosuchus | G. carolinensis | Dorchester County, South Carolina.[15] | ChM PV 4282 (mandible).[15] | Now moved to the genus Thecachampsa. | |
Thecachampsa | T. carolinensis | Dorchester County, South Carolina.[15] | ChM PV 4282 (mandible).[15] | A gavialoid. |
Turtles | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Ashleychelys | A. palmeri | East bank of Limehouse Branch, Berkeley County, South Carolina.[16] | A cheloniid also found in the Chandler Bridge Formation. | ||
Carolinochelys | C. wilsoni | MCZ 1005-A (a nearly complete skull) and ChM PV8309.[16] | A cheloniid also found in the Chandler Bridge Formation. | ||
Natemys | N. sp. 1 | Givhan's Ferry Member.[17] | CCNHM 4288 (a non-ridged carapacial ossicle).[17] | A dermochelyid also found in the Chandler Bridge Formation. | |
N. sp. 2 | ?Givhan's Ferry Member and ?Runnymede Marl Member.[17] | CCNHM 4287.1 and 4287.2 (a pair of associated non-ridged carapacial ossicles) and CCNHM
4910 (a non-ridged ossicle).[17] |
A dermochelyid. | ||
Procolpochelys | P. charlestonensis | A humerus (MCZ 1005-B).[16] | A cheloniid also found in the Chandler Bridge Formation. | ||
cf. Psephophorus | cf. P. sp. | Givhan's Ferry Member.[17] | CCNHM 5460 (an isolated non-ridged carapacial ossicle).[17] | A dermochelyid also found in the Chandler Bridge Formation. |
Fish
editCartilaginous fish
editRays | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Dasyatis | "D." sp. | Givhans Ferry Member.[1] | Multiple teeth.[1] | A stingray. | |
Mobula | "M." sp. | Givhans Ferry Member.[1] | A devil ray. | ||
Plinthicus | P. sp. | Givhans Ferry Member.[1] | A tooth (SC2007.36.48).[1] | ||
Raja | R. mccollumi | Givhans Ferry Member.[1] | Abundant teeth.[1] | A skate also found in the Chandler Bridge Formation. | |
"R." sp. | Givhans Ferry Member.[1] | Multiple teeth.[1] | A skate, appears to be conspecific with R. sp. from the Chandler Bridge Formation. | ||
Rhinoptera | "R." sp. | Givhans Ferry Member.[1] | A broken symphyseal tooth (SC2015.29.30).[1] | A cownose ray. | |
Rhynchobatus | R. sp. | Givhans Ferry Member.[1] | Teeth (SC2007.36.39, SC2015.29.13, SC2015.29.32).[1] | A wedgefish. | |
Taeniurops | "T." cavernosus | Givhans Ferry Member.[1] | Multiple teeth.[1] | A stingray. |
Sharks | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Araloselachus | A. sp. | May be derived from the Givhans Ferry Member.[1] | Teeth.[1] | ||
Carcharhinus | C. gibbesi | Givhans Ferry Member.[1] | A large number of teeth.[1] | The most common shark in the Ashley Formation sample. | |
Galeocerdo | G. aduncus | Givhans Ferry Member.[1] | Teeth.[1] | A ground shark. | |
Ginglymostomatidae | Gen. et. sp. indeterminate | Givhans Ferry Member.[1] | A partial tooth (SC2007.36.208).[1] | A nurse shark. | |
Hemipristis | H. cf. H. serra | Givhans Ferry Member.[1] | 4 teeth (SC2007.36.7, SC2007.36.8, SC2007.36.9).[1] | A weasel shark. | |
Otodus | O. angustidens | Ashley River, in the Givhans Ferry Member.[1] | Ablated specimens.[1] | A megatoothed shark. | |
Pachyscyllium | P. sp. | Givhans Ferry Member.[1] | A tooth (SC2007.36.5).[1] | A catshark. | |
Physogaleus | P. cf. P. contortus | Givhans Ferry Member.[1] | SC2007.36.19, SC2007.36.20 (four teeth), SC2015.29.27.[1] | A ground shark. | |
P. sp. | Givhans Ferry Member.[1] | SC2007.36.21 and SC2007.36.22.[1] | A ground shark. | ||
Pristiophorus | P. sp. | Givhans Ferry Member.[1] | An incomplete tooth crown (SC2015.29.20).[1] | A sawshark. | |
Scyliorhinus | S. weemsi | Givhans Ferry Member.[1] | Multiple teeth.[1] | A catshark. | |
Sphyrnidae | Gen. et. sp. indeterminate | Givhans Ferry Member.[1] | SC2007.36.23, SC2007.36.24, SC2007.36.25 (11 teeth), SC2007.36.26, SC2007.36.27, SC2007.36.28 (five teeth).[1] | A hammerhead shark. | |
Squalus | S. sp. | Givhans Ferry Member.[1] | Teeth.[1] | A spurdog. | |
Squatina | S. sp. | Givhans Ferry Member.[1] | Teeth (SC2007.36.4, SC2007.36.126, SC2007.36.149, SC2007.36.227, SC2015.29.18, SC2015.29.19, SC2015.33.2).[1] | An angelshark. | |
Trigonotodus | T. alteri | Teeth.[1] | A thresher shark. |
Bony fish
editBony fish | |||||
---|---|---|---|---|---|
Genus | Species | Presence | Material | Notes | Images |
Albula | A. sp. | Givhans Ferry Member.[1] | SC2007.36.165, SC2007.36.228 (five teeth), SC2015.29.200 (66 teeth), SC2015.29.201, SC2015.29.204.[1] | A bonefish. | |
Archosargus | A. sp. | Givhans Ferry Member.[1] | Multiple teeth.[1] | A sea bream. | |
?Calamus | ?C. sp. | Givhans Ferry Member.[1] | Teeth.[1] | A sea bream. | |
Diplodus | D. sp. | Givhans Ferry Member.[1] | SC2015.29.195 (13 teeth), SC2015.29.222, SC2015.29.223, SC2015.29.224.[1] | A sea bream. | |
?Labridae | Gen. et. sp. indet. | Givhans Ferry Member.[1] | Multiple teeth.[1] | A wrasse. | |
Osteoglossidae | Gen. et. sp. indeterminate | Givhans Ferry Member.[1] | Teeth.[1] | ||
Palaeocybium | P. sp. | Givhans Ferry Member.[1] | A tooth (SC2007.36.122), a neurocranium and a left dentary.[1] | A scombrid. | |
Paralichthyidae | Gen. et. sp. indeterminate | Givhans Ferry Member.[1] | A tooth (SC2015.36.259).[1] | A flatfish. | |
?Pogonias | ?P. sp. | Givhans Ferry Member.[1] | Teeth.[1] | A croaker. | |
Scomberomorus | S. sp. | Givhans Ferry Member.[1] | Teeth.[1] | A scombrid. | |
?Sciaenops | ?S. sp. | Givhans Ferry Member.[1] | Many teeth.[1] | A croaker. | |
Sphyraena | S. sp. | Givhans Ferry Member.[1] | Many cheek and laniary teeth.[1] | A barracuda. | |
Trichiurides | T. cf. T. sagittidens | Givhans Ferry Member.[1] | 2 ablated laniary teeth.[1] | A cutlassfish. |
See also
editReferences
edit- ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs Cicimurri, David J.; Knight, James L.; Ebersole, Jun A. (2022). "Early Oligocene (Rupelian) fishes (Chondrichthyes, Osteichthyes) from the Ashley Formation (Cooper Group) of South Carolina, USA". PaleoBios. 39 (1). doi:10.5070/P939056976. ISSN 0031-0298. S2CID 247912932.
- ^ J., Ray, Clayton Edward. Emry, Robert (2002). Cenozoic mammals of land and sea : tributes to the career of Clayton E. Ray. Smithsonian Institution Press. pp. 179–183. OCLC 1035595001.
{{cite book}}
: CS1 maint: multiple names: authors list (link) - ^ a b Godfrey, Stephen J.; Uhen, Mark D.; Osborne, Jason E.; Edwards, Lucy E. (January 2016). "A new specimen of Agorophius pygmaeus (Agorophiidae, Odontoceti, Cetacea) from the early Oligocene Ashley Formation of South Carolina, USA". Journal of Paleontology. 90 (1): 154–169. doi:10.1017/jpa.2016.4. ISSN 0022-3360. S2CID 132593720.
- ^ a b Boessenecker, Robert W.; Ahmed, Erum; Geisler, Jonathan H. (2017-11-08). "New records of the dolphin Albertocetus meffordorum (Odontoceti: Xenorophidae) from the lower Oligocene of South Carolina: Encephalization, sensory anatomy, postcranial morphology, and ontogeny of early odontocetes". PLOS ONE. 12 (11): e0186476. Bibcode:2017PLoSO..1286476B. doi:10.1371/journal.pone.0186476. ISSN 1932-6203. PMC 5695589. PMID 29117197.
- ^ Boessenecker, Robert W.; Churchill, Morgan; Buchholtz, Emily A.; Beatty, Brian L.; Geisler, Jonathan H. (2020-08-17). "Convergent Evolution of Swimming Adaptations in Modern Whales Revealed by a Large Macrophagous Dolphin from the Oligocene of South Carolina". Current Biology. 30 (16): 3267–3273.e2. doi:10.1016/j.cub.2020.06.012. ISSN 0960-9822. PMID 32649912. S2CID 220435400.
- ^ a b Sanders, Albert E.; Geisler, Jonathan H. (2015-01-02). "A new basal odontocete from the upper Rupelian of South Carolina, U.S.A., with contributions to the systematics of Xenorophus and Mirocetus (Mammalia, Cetacea)". Journal of Vertebrate Paleontology. 35 (1): e890107. doi:10.1080/02724634.2014.890107. ISSN 0272-4634. S2CID 85040115.
- ^ Geisler, Jonathan H.; Boessenecker, Robert W.; Brown, Mace; Beatty, Brian L. (2017-07-10). "The Origin of Filter Feeding in Whales". Current Biology. 27 (13): 2036–2042.e2. doi:10.1016/j.cub.2017.06.003. ISSN 0960-9822. PMID 28669761. S2CID 36765725.
- ^ Albright, L. Barry; Sanders, Albert E.; Geisler, Jonathan H. (April 2019). "An Unexpectedly Derived Odontocete from the Ashley Formation (Upper Rupelian) of South Carolina, U.S.A." Journal of Vertebrate Paleontology. 38 (4): (1)-(15). doi:10.1080/02724634.2018.1482555. ISSN 0272-4634. S2CID 92830510.
- ^ Sanders Albert, E.; Barnes Lawrence, G. (2002-11-21). "Paleontology of the late Oligocene Ashley and Chandler Bridge Formations of South Carolina; 2, Micromysticetus rothauseni, a primitive cetoteriid mysticete (Mammalia, Cetacea)". Smithsonian Contributions to Paleobiology. 93: 271–293.
- ^ L. Agassiz. 1848. [on the species Saurocetus gibbesii]. Proceedings of the Academy of Natural Sciences of Philadelphia 4:4-5
- ^ Dooley, Alton C. Jr (2003). "A Review of the Eastern North American Squalodontidae (Mammalia: Cetacea)". Jeffersoniana (11): 1–26.
- ^ Kellogg, Remington (1923). "Description of an apparently new toothed cetacean from South Carolina". Smithsonian Miscellaneous Collections. 76 (7): 1–7.
- ^ Domning, Daryl P. (1997-06-19). "Fossil Sirenia of the west Atlantic and Caribbean region. VI. Crenatosiren olseni (Reinhart, 1976)". Journal of Vertebrate Paleontology. 17 (2): 397–412. doi:10.1080/02724634.1997.10010984. ISSN 0272-4634.
- ^ Domning, Daryl P.; Beatty, Brian L. (2019-05-04). "Fossil Sirenia of the West Atlantic and Caribbean region. XII. Stegosiren macei, gen. et sp. nov". Journal of Vertebrate Paleontology. 39 (3): e1650369. doi:10.1080/02724634.2019.1650369. ISSN 0272-4634. S2CID 203407242.
- ^ a b c d R.., Erickson, Bruce (1996). The estuarine crocodile Gavialosuchus Carolinensis n.sp. (Crocodylia : Eusuchia) from the late Oligocene of South Carolina, North America (PDF). Science Museum of Minnesota. OCLC 491733633.
{{cite book}}
: CS1 maint: multiple names: authors list (link) - ^ a b c Weems, Robert E.; Sanders, Albert E. (January 2014). "Oligocene pancheloniid sea turtles from the vicinity of Charleston, South Carolina, U.S.A." Journal of Vertebrate Paleontology. 34 (1): 80–99. doi:10.1080/02724634.2013.792826. ISSN 0272-4634. S2CID 128543145.
- ^ a b c d e f "Multispecies leatherback turtle assemblage from the Oligocene Chandler Bridge and Ashley formations of South Carolina, USA - Acta Palaeontologica Polonica". www.app.pan.pl. Retrieved 2022-09-11.
- Various Contributors to the Paleobiology Database. "Fossilworks: Gateway to the Paleobiology Database". Retrieved 17 December 2021.