Myriopteris scabra, commonly known as rough lipfern, is a species of cheilanthoid fern native to Mexico and the south-western United States (Texas).[2]
| Myriopteris scabra | |
|---|---|
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Scientific classification 
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| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Division: | Polypodiophyta |
| Class: | Polypodiopsida |
| Order: | Polypodiales |
| Family: | Pteridaceae |
| Genus: | Myriopteris |
| Species: | M. scabra
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| Binomial name | |
| Myriopteris scabra (C.Chr.) Grusz & Windham
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| Synonyms[1] | |
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Description
editMyriopteris scabra grows from a short creeping rhizome, usually 4–7 mm in diameter with brown scales. The leaves (fronds) are clustered and may range greatly in size from 5–30 cm long. The leaf petiole is black to dark brown. The leaf blade is 1–4 cm wide, linear-oblong to lanceolate, and up to pinnate-pinnatifid to 2-pinnate. The rachis has scattered linear-lanceolate scales and dimorphic pubescence, abaxially sparsely hirsute, adaxially covered with tortuous appressed hairs. The ultimate leaflets are narrowly elliptic to elongate-deltate, not beadlike, and up to 3–5 mm long. The upper leaflet surface has a distinctive rough or spiky surface, which distinguishes this species from most other Myriopteris, and gives rise to the specific name scabra (Latin for rough or coarse). The leaflet edges are only folded under at their margins, barely concealing the sori (when present), which form a continuous bead around leaflet margins.[3]
Range and habitat
editMyriopteris scabra is native to northern Mexico, the Yucatan Peninsula, and to central and western Texas, extending north into Oklahoma. It grows on rocks and in rock crevices, mostly restricted to limestone, at elevations from 100 to 1400 meters.[3] Some close relatives of Myriopteris scabra are also calcareous soil/limestone specialists, including Myriopteris alabamensis, Myriopteris aemula, and Myriopteris microphylla.[4][5] These species have ranges in the southern United States, Mexico, and Caribbean islands. Interestingly, another Myriopteris limestone specialist Myriopteris gracilis is not closely related to Myriopteris scabra. M. gracilis has a range that includes high mountains in the southwest United States and extends to much colder regions including Montana, Idaho, and British Columbia.[6]
Taxonomy
editAccording to plastid DNA sequence analysis, Myriopteris scabra is in the alabamensis clade of Myriopteris and its closest relative is Myriopteris fimbriata.[7]
References
edit- ^ Grusz & Windham 2013.
- ^ "Myriopteris scabra (Rough Lipfern)". iNaturalist. Retrieved 2022-10-26.
- ^ a b SEINet: Myriopteris scabra
- ^ SEINet: Myriopteris aemula
- ^ SEINet: Myriopteris microphylla
- ^ "The Jepson Herbarium".
- ^ Grusz et al. 2014.
Works cited
edit- Grusz, Amanda L.; Windham, Michael D. (2013). "Toward a monophyletic Cheilanthes: The resurrection and recircumscription of Myriopteris (Pteridaceae)". PhytoKeys (32): 49–64. doi:10.3897/phytokeys.32.6733. PMC 3881352. PMID 24399906.
- Grusz, Amanda L.; Windham, Michael D.; Yatskievych, George; Huiet, Lane; Gastony, Gerald J.; Pryer, Kathleen M. (2014). "Patterns of Diversification in the Xeric-adapted Fern Genus Myriopteris (Pteridaceae)" (PDF). Systematic Botany. 39 (3): 698–714. doi:10.1600/036364414X681518. JSTOR 24546228. S2CID 16969741.