Orthogonius

Orthogonius is a genus of beetles in the family Carabidae, containing the following species:

• Orthogonius aberlenci Tian & Deuve, 2016 (Laos)
• Orthogonius acrogonus (Wiedemann, 1819) (Malaysia, Indonesia, and Borneo)
• Orthogonius acutangulus Chaudoir, 1878 (Sri Lanka)
• Orthogonius adoriae Tian & Deuve, 2006 (India)
• Orthogonius aemulus Péringuey, 1896 (Zimbabwe, Botswana, and Namibia)
• Orthogonius alternans (Wiedemann, 1823) (Indomalaya)
• Orthogonius alutaceus Quedenfeldt, 1883 (Africa)
• Orthogonius andrewesi Emden, 1928 (Philippines)
• Orthogonius andrewesianus Tian & Deuve, 2006 (Malaysia, Singapore, and Indonesia)
• Orthogonius angkor Tian & Deuve, 2006 (Cambodia)
• Orthogonius angkorensis Tian & Deuve, 2008 (Cambodia)
• Orthogonius angustus Chaudoir, 1872 (Thailand, Vietnam, and Malaysia)
• Orthogonius annamensis Jedlicka, 1965 (Vietnam)
• Orthogonius annamicola Tian & Deuve, 2006 (Vietnam)
• Orthogonius annemariae Tian & Deuve, 2006 (India)
• Orthogonius assamensis Tian & Deuve, 2006 (India)
• Orthogonius assamicola Tian & Deuve, 2006 (India)
• Orthogonius assmuthi Wasmann, 1920 (India)
• Orthogonius baconi Chaudoir, 1872 (India and Myanmar)
• Orthogonius baconioides Tian & Sabu, 2009 (India)
• Orthogonius balthasari Jedlicka, 1935 (Philippines)
• Orthogonius batesi Tian & Deuve, 2003 (Sri Lanka)
• Orthogonius batesianus Tian & Deuve, 2007
• Orthogonius batesioides Tian & Deuve, 2006 (India)
• Orthogonius baumi Jedlicka, 1955 (Malaysia)
• Orthogonius becqueti Burgeon, 1937 (Democratic Republic of the Congo and Burundi)
• Orthogonius bellus Tian & Deuve, 2016 (Vietnam)
• Orthogonius bicolor Jedlicka, 1965 (Vietnam)
• Orthogonius borneoensis Tian & Deuve, 2006 (Indonesia and Borneo)
• Orthogonius brancuccii Tian & Deuve, 2006 (Thailand)
• Orthogonius brevilabris Kolbe, 1889 (Democratic Republic of the Congo and Angola)
• Orthogonius brevithorax Dejean, 1825 (Africa)
• Orthogonius bruggei Tian & Deuve, 2006 (Malaysia)
• Orthogonius buquetii Chaudoir, 1850 (Sierra Leone and Ivory Coast)
• Orthogonius burckhardti Tian & Deuve, 2006
• Orthogonius burmanensis Tian & Deuve, 2006 (Myanmar)
• Orthogonius caffer Boheman, 1848 (Zimbabwe and South Africa)
• Orthogonius cambodgensis Tian & Deuve, 2006 (Cambodia)
• Orthogonius canaraensis Tian & Deuve, 2006 (India)
• Orthogonius canaraicola Tian & Deuve, 2006 (India)
• Orthogonius capucinus Boheman, 1848 (Zimbabwe and South Africa)
• Orthogonius carbonicolor Tian & Deuve, 2006 (Myanmar)
• Orthogonius carinatus Tian & Deuve, 2013 (Laos)
• Orthogonius celebesicus Tian & Deuve, 2006 (Indonesia)
• Orthogonius ceylanicus Tian & Deuve, 2016 (Sri Lanka)
• Orthogonius chaudoiri Tian & Deuve, 2001 (Vietnam)
• Orthogonius cheni Tian & Deuve, 2001 (China)
• Orthogonius chiangensis Tian & Deuve, 2006 (Thailand)
• Orthogonius clarkii Murray, 1858 (Africa)
• Orthogonius confusus Tian & Deuve, 2006 (Malaysia)
• Orthogonius constanti Tian & Deuve, 2006 (Cambodia)
• Orthogonius constrictus Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius coomani Tian & Deuve, 2006 (Vietnam)
• Orthogonius coomanioides Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius coracinus Kolbe, 1895 (Uganda, Rwanda, Burundi, and Tanzania)
• Orthogonius crassicrus Chaudoir, 1872 (Indonesia and Borneo)
• Orthogonius cruralis Putzeys in Chaudoir, 1872
• Orthogonius curvatus Tian & Deuve, 2006 (India)
• Orthogonius cyclothorax Tian & Deuve, 2007 (Singapore)
• Orthogonius davidi Chaudoir, 1878 (China and Taiwan)
• Orthogonius deletus Schmidt-Goebel, 1846 (China, India, Southeast Asia)
• Orthogonius deuvei Tian & Kirschenhofer, 2013 (Vietnam)
• Orthogonius dispar Bates, 1892 (Myanmar)
• Orthogonius dohertyi Tian & Deuve, 2006 (Malaysia)
• Orthogonius dongnanya Tian & Deuve, 2006 (Singapore, Indonesia, and Borneo)
• Orthogonius doriae Putzeys in Chaudoir, 1872 (Indonesia and Borneo)
• Orthogonius drescheri Liebke, 1937 (Indonesia)
• Orthogonius drumonti Tian & Deuve, 2006 (Indonesia)
• Orthogonius duboisi Tian & Deuve, 2006 (China)
• Orthogonius duplicatus (Wiedemann, 1819) (Indomalaya)
• Orthogonius dupuisi Basilewsky, 1948 (Zambia)
• Orthogonius dupuisianus Tian & Deuve, 2010 (Indonesia)
• Orthogonius dureli Tian & Deuve, 2005 (Bhutan and India)
• Orthogonius edentatus Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
• Orthogonius elisabethanus Burgeon, 1937 (Democratic Republic of the Congo)
• Orthogonius emarginatus Tian & Deuve, 2007 (Singapore)
• Orthogonius equimarginalis Tian & Deuve, 2006 (Indonesia)
• Orthogonius euthyphallus Tian & Deuve, 2013 (Laos and Vietnam)
• Orthogonius fairmairei Tian & Deuve, 2006 (Singapore)
• Orthogonius feai Tian & Deuve, 2007
• Orthogonius femoralis Chaudoir, 1848 (Sri Lanka and India)
• Orthogonius flavipes Deuve, 2004 (Indonesia)
• Orthogonius flavus Jedlicka, 1964 (India)
• Orthogonius foveiclypeus Tian & Deuve, 2004 (Indonesia)
• Orthogonius fryi Tian & Deuve, 2006 (Myanmar)
• Orthogonius fugax Chaudoir, 1872 (Sri Lanka)
• Orthogonius gracililamella Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius gracilipes Tian & Deuve, 2006 (India)
• Orthogonius gracilis Tian & Deuve, 2006 (India)
• Orthogonius grootaerti Tian & Deuve, 2006 (Thailand)
• Orthogonius hageni Oberthür, 1883 (Indonesia)
• Orthogonius himalaya Tian & Deuve, 2006 (Bhutan and India)
• Orthogonius himalayicus Tian & Deuve, 2005 (Bhutan and India)
• Orthogonius hirtulus Tian & Deuve, 2006 (Indonesia)
• Orthogonius hirtus Chaudoir, 1872 (Malaysia and Indonesia)
• Orthogonius hopei Gray, 1832 (Indomalaya)
• Orthogonius huananoides Tian & Deuve, 2006 (Thailand and Vietnam)
• Orthogonius huananus Tian & Deuve, 2001 (China)
• Orthogonius hypocrita Chaudoir, 1872 (Indonesia and Philippines)
• Orthogonius hypocritoides Jedlicka, 1935 (Philippines)
• Orthogonius impunctipennis Quedenfeldt, 1883 (Democratic Republic of the Congo and Angola)
• Orthogonius indicus Tian & Deuve, 2006 (India)
• Orthogonius indophilus Tian & Deuve, 2006 (India)
• Orthogonius inexpectatus Tian & Deuve, 2006 (India)
• Orthogonius inops Chaudoir, 1872 (Indonesia)
• Orthogonius insularis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
• Orthogonius intermedius Chaudoir, 1872 (Indonesia)
• Orthogonius javanus Tian & Deuve, 2006 (Taiwan and Indonesia)
• Orthogonius jianfengling Tian & Deuve, 2001 (China)
• Orthogonius kalimantanensis Tian & Deuve, 2006 (Indonesia and Borneo)
• Orthogonius kapiriensis Burgeon, 1937 (Democratic Republic of the Congo)
• Orthogonius katangensis Burgeon, 1937 (Democratic Republic of the Congo)
• Orthogonius kickeli Kolbe, 1895 (Namibia)
• Orthogonius kirirom Tian & Deuve, 2008 (Thailand and Cambodia)
• Orthogonius klickai Jedlicka, 1935 (Philippines)
• Orthogonius kubani Tian & Deuve, 2006 (Thailand)
• Orthogonius kumatai Habu, 1979 (Nepal)
• Orthogonius lancangjiang Tian & Deuve, 2006 (Laos)
• Orthogonius lao Tian & Deuve, 2006 (Laos)
• Orthogonius latreillei Tian & Deuve, 2006 (India)
• Orthogonius latus Hope, 1842 (Africa)
• Orthogonius ledouxi Tian & Deuve, 2006 (Vietnam)
• Orthogonius legrandi Tian & Deuve, 2006 (Malaysia)
• Orthogonius lieftincki Andrewes, 1936 (Indonesia)
• Orthogonius limbourgi Tian & Deuve, 2016 (Vietnam)
• Orthogonius loeicus Tian & Deuve, 2006 (Thailand)
• Orthogonius loeiensis Tian & Deuve, 2006 (Thailand)
• Orthogonius longicornis Chaudoir, 1872 (Sri Lanka and Thailand)
• Orthogonius longilamella Tian & Deuve, 2008
• Orthogonius longipennis Hope, 1842
• Orthogonius longiphallus Tian & Deuve, 2005 (Bhutan)
• Orthogonius lucidus Bates, 1891 (India)
• Orthogonius luzonicus Chaudoir, 1872 (Philippines)
• Orthogonius macrophthalmus Tian & Deuve, 2013 (Vietnam)
• Orthogonius maindronianus Tian & Deuve, 2006 (India)
• Orthogonius makiling Tian & Deuve, 2006 (Philippines)
• Orthogonius malacca Tian & Deuve, 2006 (Malaysia)
• Orthogonius malaisei Andrewes, 1947 (Myanmar)
• Orthogonius malaya Tian & Deuve, 2006 (Malaysia)
• Orthogonius malaysiaensis Tian & Deuve, 2006 (Malaysia)
• Orthogonius medanensis Tian & Deuve, 2006 (Indonesia)
• Orthogonius meghalayaensis Tian & Deuve, 2016 (India)
• Orthogonius mekongensis Tian & Deuve, 2006 (Laos)
• Orthogonius melanipes Tian & Deuve, 2006 (India)
• Orthogonius mellyi (Chaudoir, 1850) (Bangladesh, India, and Myanmar)
• Orthogonius minimus Tian & Deuve, 2006 (India)
• Orthogonius mniszechi Chaudoir, 1872 (India, Malaysia, Indonesia, and Borneo)
• Orthogonius moestus Chaudoir, 1872
• Orthogonius monolophus Andrewes, 1931 (Indonesia and Borneo)
• Orthogonius montanus Tian & Deuve, 2006 (Myanmar)
• Orthogonius morvani Tian & Deuve, 2003 (Thailand)
• Orthogonius morvanianus Tian & Deuve, 2016 (Thailand)
• Orthogonius mouhoti Chaudoir, 1872 (Southeast Asia)
• Orthogonius myanmarensis Tian & Deuve, 2006 (Myanmar)
• Orthogonius nahaeo Tian & Deuve, 2006 (Thailand)
• Orthogonius nepalensis Habu, 1979 (Nepal)
• Orthogonius nepalicus Tian & Deuve, 2006 (Nepal)
• Orthogonius niger Jedlicka, 1935 (Philippines)
• Orthogonius nigripes Tian & Deuve, 2001 (China and Vietnam)
• Orthogonius nongfaensis Tian & Kirschenhofer, 2013 (Laos)
• Orthogonius nyassicus Kolbe, 1895
• Orthogonius obliquepes Tian & Deuve, 2006 (Philippines)
• Orthogonius opacus Schmidt-Goebel, 1846 (Indomalaya)
• Orthogonius orphnodes Andrewes, 1930 (Indonesia)
• Orthogonius ovatulus Tian & Deuve, 2003 (Sri Lanka)
• Orthogonius ovatus Chaudoir, 1878 (Indonesia)
• Orthogonius pachlatkoi Tian & Deuve, 2006 (Thailand)
• Orthogonius palangbangensis Tian & Deuve, 2006 (Indonesia)
• Orthogonius panghongae Tian & Deuve, 2006 (China)
• Orthogonius pangi Tian & Deuve, 2006 (Thailand and Vietnam)
• Orthogonius parallelus Chaudoir, 1872 (Sri Lanka)
• Orthogonius parastantoni Tian & Deuve, 2006 (Sri Lanka)
• Orthogonius parasuturalis Tian & Deuve, 2006 (Indonesia)
• Orthogonius parcepunctatus Kolbe, 1895 (Tanzania)
• Orthogonius paris Tian & Deuve, 2006 (Thailand)
• Orthogonius parvus Chaudoir, 1872 (India)
• Orthogonius pectinatus Tian & Deuve, 2016 (Thailand and Vietnam)
• Orthogonius perakensis Tian & Deuve, 2006 (Malaysia)
• Orthogonius perakicus Tian & Deuve, 2007 (Malaysia)
• Orthogonius perpuncticollis Burgeon, 1937 (Africa)
• Orthogonius petiolaris Tian & Deuve, 2006 (India)
• Orthogonius piceus Chaudoir, 1872 (Malaysia and Indonesia)
• Orthogonius picilabris (W.S.MacLeay, 1825) (Malaysia, Indonesia, and Borneo)
• Orthogonius picipennis Chaudoir, 1872 (Cambodia and Vietnam)
• Orthogonius pinguis Murray, 1858 (Nigeria)
• Orthogonius pinophilus Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius planiger (Walker, 1858) (Sri Lanka)
• Orthogonius plebius Tian & Deuve, 2006 (Sri Lanka)
• Orthogonius poggii Tian & Deuve, 2007
• Orthogonius politior Tian & Deuve, 2016 (Laos)
• Orthogonius politus Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
• Orthogonius pollinctor Basilewsky, 1948 (Democratic Republic of the Congo)
• Orthogonius pradieri Chaudoir, 1872
• Orthogonius pseudochaudoiri Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius pseudocheni Tian & Deuve, 2006 (Vietnam)
• Orthogonius pseudolongicornis Tian & Deuve, 2006 (Southeast Asia)
• Orthogonius pseudomouhoti Tian & Deuve, 2008 (Cambodia)
• Orthogonius punctatellus Tian & Deuve, 2006 (Indonesia)
• Orthogonius punctulatus Chaudoir, 1872 (India)
• Orthogonius punctum Tian & Deuve, 2016 (Thailand and Vietnam)
• Orthogonius putzeysi Tian & Deuve, 2006 (Sri Lanka)
• Orthogonius quadrisetosus Tian & Deuve, 2006 (Malaysia)
• Orthogonius rhodesianus Csiki, 1932
• Orthogonius rhodesiensis Lorenz, 1998 (Zambia)
• Orthogonius ribbei Tian & Deuve, 2003 (Indonesia)
• Orthogonius rogueti Tian & Deuve, 2016 (India)
• Orthogonius rotundatus Tian & Deuve, 2006
• Orthogonius rufiventris Bates, 1892 (Myanmar)
• Orthogonius rufotestaceus G.Müller, 1941 (Ethiopia)
• Orthogonius sabahensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
• Orthogonius sabahicus Tian & Deuve, 2016 (Malaysia, Indonesia, and Borneo)
• Orthogonius salvazai Tian & Deuve, 2006 (Laos)
• Orthogonius sarawakensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
• Orthogonius saundersi Andrewes, 1926 (Malaysia and Singapore)
• Orthogonius schaumi Chaudoir, 1872 (Sri Lanka and Indonesia)
• Orthogonius schaumioides Tian & Deuve, 2006 (India)
• Orthogonius schauteni Tian & Deuve, 2006 (Malaysia)
• Orthogonius schmidtgoebeli Chaudoir, 1872 (Myanmar)
• Orthogonius senegalensis Dejean, 1831 (Africa)
• Orthogonius setosopalpiger Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius siamensis Tian & Deuve, 2006 (Thailand)
• Orthogonius similaris Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius simplicatus Tian & Deuve, 2006 (Indonesia)
• Orthogonius singaporensis Tian & Deuve, 2006 (Singapore)
• Orthogonius sinuatiphallus Tian & Deuve, 2001 (China)
• Orthogonius smetsi Tian & Deuve, 2006 (Cambodia)
• Orthogonius solidicornis Tian & Deuve, 2003 (China)
• Orthogonius solidus Tian & Deuve, 2006 (India)
• Orthogonius spinosus Burgeon, 1937 (Africa)
• Orthogonius srilankaicus Tian & Deuve, 2003 (Sri Lanka)
• Orthogonius stantoni Tian & Deuve, 2006 (India)
• Orthogonius sterbai Jedlicka, 1935 (Philippines)
• Orthogonius stygius Andrewes, 1930 (Malaysia and Indonesia)
• Orthogonius sulawesicus Tian & Deuve, 2003 (Indonesia)
• Orthogonius sulcatoides Tian & Deuve, 2006 (India, Myanmar, and Malaysia)
• Orthogonius sulcatus Schmidt-Goebel, 1846 (Myanmar and Cambodia)
• Orthogonius sumatraicus Tian & Deuve, 2006 (Indonesia)
• Orthogonius suturalis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
• Orthogonius taghavianae Tian; Deuve & Felix, 2012 (Thailand)
• Orthogonius taiwanicus Tian & Deuve, 2006 (Taiwan)
• Orthogonius tangsen Tian & Deuve, 2006 (India)
• Orthogonius tenasserimensis Tian & Deuve, 2006 (Myanmar)
• Orthogonius termiticola Wasmann, 1902 (Malaysia)
• Orthogonius thaicus Tian & Deuve, 2003 (Thailand)
• Orthogonius thaiensis Tian & Deuve, 2006 (Thailand)
• Orthogonius thailandensis Tian & Deuve, 2006 (Thailand)
• Orthogonius thoracicus Gestro, 1875
• Orthogonius tikekee Tian & Deuve, 2006 (Myanmar)
• Orthogonius tonkinensis Tian & Deuve, 2006 (Vietnam)
• Orthogonius tonkinicus Tian & Deuve, 2006 (Vietnam)
• Orthogonius travancore Tian & Deuve, 2006 (India)
• Orthogonius tricarinatus Burgeon, 1937 (Africa)
• Orthogonius uncipennis Tian & Deuve, 2006 (Nepal)
• Orthogonius vari Tian; Deuve & Felix, 2012 (Thailand and Cambodia)
• Orthogonius variabilis Tian; Deuve & Felix, 2012 (China and Thailand)
• Orthogonius virgulatus Andrewes, 1931 (Indonesia and Borneo)
• Orthogonius wallardiensis Tian & Deuve, 2006 (India)
• Orthogonius wrasei Tian & Deuve, 2016 (Myanmar)
• Orthogonius xanthomerus L.Redtenbacher, 1868 (China)
• Orthogonius yoga Tian & Deuve, 2006 (India)
• Orthogonius yunnanensis Tian & Deuve, 2001 (China)

Orthogonius
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Superfamily:	Caraboidea
Family:	Carabidae
Subfamily:	Orthogoniinae
Genus:	Orthogonius W.S.MacLeay, 1825