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Paraná Formation

The Paraná Formation is a geologic formation in Argentina. It preserves fossils dating back to the late Miocene epoch.

Paraná Formation
Stratigraphic range: Late Miocene
TypeGeological formation
Location
RegionEntre Ríos Province
Country Argentina
Type section
Named forParaná Basin

Fossil content

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Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Mammals

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Cetaceans

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Cetaceans reported from the Paraná Formation
Genus Species Presence Material Notes Images
cf. Aulophyseter cf. A. sp. Cantera del Puerto Viejo near Paraná, Entre Ríos.[1] Incomplete tooth (MACN Pv 8926).[1] Now deemed an indeterminate physeteroid.[2]
Balaenidae gen. et. sp. indet. Indeterminate Right tympanic bulla (MACN Pv 13221).[1] A balaenid.
Balaenoptera B. sp. Right tympanic bulla (MACN Pv 13220).[1] A rorqual.
Balaenopteridae indet. South Bank of Ensenada Creek.[3] Cubitus & radius (CICYTTP-PV-1-197) bearing bite marks matching Cosmopolitodus hastalis.[3] A rorqual.
Physeteroidea gen. et. sp. indet. Indeterminate Cantera del Puerto Viejo near Paraná, Entre Ríos.[2] An almost complete tooth (MAS Pv 1361) & a fragmentary tooth (MACN Pv 8926).[2] Previously identified as cf. Aulophyseter sp.[2]
Platanistoidea indet. Left tympanic bulla (MACN Pv 15997).[1] A relative of river dolphins.
Pontistes P. rectifrons [4] A pontoporiid.
Prionodelphis P. rovereti A single tooth.[5] A dubious delphinoid, other specimens once referred to this taxon are now thought to represent monachine seals.[5]

Pinnipeds

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Pinnipeds reported from the Paraná Formation
Genus Species Presence Material Notes Images
Properiptychus P. argentinus Jaw & limb elements.[6] A monachine seal.

Rodents

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Rodents reported from the Paraná Formation
Genus Species Presence Material Notes Images
cf. Cardiatherium cf. C. sp. indet. La Juanita locality.[7] Left isolated cheek tooth (CICYTTP-PV-M-1-147) & right upper cheek tooth (CICYTTP-PV-M-1-148).[7] A hydrochoerine.
 
Caviidae gen. & sp. indet. Indeterminate La Juanita locality.[7] A left upper M1-2 (CICYTTP-PV-M-2-267).[7] A cavy.
Plesiacarechimys P. nov. sp.? La Juanita locality.[7] Right lower molar (CICYTTP-PV-M-1-145).[7] An octodontoid.

Sirenians

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Sirenians reported from the Paraná Formation
Genus Species Presence Material Notes Images
Dioplotherium D. sp. Cliffs along Paraná River, Villa Urquiza.[8] Upper left molar (CORD-PZ 4301).[8] A dugongid.
 
Metaxytherium M. sp. Cerro La Matanza locality, Victoria City.[8] Posterior end of both maxillae (MASP 373).[8] A dugongid.
 

Xenarthrans

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Xenarthrans reported from the Paraná Formation
Genus Species Presence Material Notes Images
Scirrotherium S. sp. indet. La Juanita locality.[7] A fixed osteoderm (CICYTTP-PV-M-1-144).[7] A pampatheriid.
 

Birds

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Birds reported from the Paraná Formation
Genus Species Presence Material Notes Images
Pelecanus P. paranensis Cerro La Matanza locality, Victoria City.[9] A nearly complete pelvis (CICYTTP-PV-A-3-277).[9] A pelican.
cf. Phoenicopterus cf. P. sp. indet. La Juanita locality.[7] Fragment of shaft with distal end of left tibiotarsus (CICYTTP-PV-A-1-12).[7] A flamingo.
 

Fish

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Bony fish

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Bony fish reported from the Paraná Formation
Genus Species Presence Material Notes Images
Siluroidae Undetermined Abundant fragmentary specimens.[10] Evidently represents extant catfish genera.[10]
Tetraodontidae indet. Indeterminate La Juanita & Toma Vieja localities.[11] Several premaxillary beaks.[11] Remains of pufferfish.

Rays

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Rays reported from the Paraná Formation
Genus Species Presence Material Notes Images
Dynatobatis D. paranensis Dermal tubercles.[10]
Myliobatis M. americanus Detached teeth.[10]
Raja R. agassizi Dermal tubercles.[10] Moved to the genus Rioraja.
Rioraja R. agassizii Dermal tubercles.[10] A skate, originally reported as Raja agassizi.
 

Sharks

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Sharks reported from the Paraná Formation
Genus Species Presence Material Notes Images
Carcharias C. (Prionodon) obliquidens Teeth.[10] Junior synonym of Squalus obliquidens.[12]
Carcharodon C. megalodon Teeth.[10] Reassigned to the genus Otodus.
 
C. sp. Arroyo Ensenada valley.[13] 2 teeth (UAP 1301 & 1303).[13] Related to the great white shark.
Cestracion C. paranensis 2 teeth.[10] Cestracion is now a junior synyonym of Heterodontus.
Cosmopolitodus C. hastalis Several teeth.[10] A mackerel shark, originally reported as Oxyrhina hastalis.
 
Galeocerdo G. aduncus Tooth.[10] A requiem shark.
 
Hemipristis H. serra A few well-preserved upper teeth.[10] A large weasel shark.
 
Heterodontus H. paranensis 2 teeth.[10] A bullhead shark, originally reported as Cestracion paranensis.
Megascyliorhinus M. trelewensis La Juanita locality.[14] 4 anterior teeth.[14] A ground shark, specimens originally labelled "Squatina".[14]
Odontaspis O. elegans Teeth.[10]
Otodus O. megalodon Teeth.[10] A megatooth shark, originally reported as Carcharodon megalodon.
 
Oxyrhina O. hastalis Several teeth.[10] Reassigned to the genus Cosmopolitodus.
Squalus S. obliquidens Teeth.[10] Originally reported as Carcharias (Prionodon) obliquidens.

Invertebrates

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Bivalves

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Bivalves reported from the Paraná Formation
Genus Species Presence Material Notes Images
Aequipecten A. paranensis Valves.[15] A scallop.
Amusium A. darwinianum [15] A scallop.
Erodona E. doellojuradoi La Juanita & La Paz.[16] 260 specimens.[16] An erodonid.
Flabellipecten F. oblongus Valves.[15] A scallop.
Polymesoda P. muravchiki La Juanita.[16] 13 specimens.[16] A cyrenid.

Bryozoans

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Bryozoans reported from the Paraná Formation
Genus Species Presence Material Notes Images
Acanthodesia A. cf. sulcata Punta Gorda, near Diamante, Entre Ríos.[17] Fragments of colonies (MLP 13471 to 13476).[17] A cheilostomatid.
Microporella M. sp. Punta Gorda, near Diamante, Entre Ríos.[17] Fragments of colonies (MLP 13477 to 13483).[17] A microporellid.
 
Schizoporella S. sp. Punta Gorda, near Diamante, Entre Ríos.[17] Fragments of colonies (MLP 13474, MLP 13484 to 13488).[17] A schizoporellid.
 
Schizoporellidae indet. Indeterminate Punta Gorda, near Diamante, Entre Ríos.[17] Fragments of colonies (MLP 13489).[17] A schizoporellid.
Schizosmittina S. sp. Punta Gorda, near Diamante, Entre Ríos.[17] Fragments of colonies (MLP 13486, MLP 13490 to 13492).[17] A bitectiporid.
Tubulipora T. sp. Punta Gorda, near Diamante, Entre Ríos.[17] Two colonies (MLP 13493 & MLP 13494).[17] A tubuliporid.
 

Echinoderms

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Echinoderms reported from the Paraná Formation
Genus Species Presence Material Notes Images
Abertella A. sp. [18] A sand dollar.
 
Amplaster A. alatus [18] A sand dollar.
A. ellipticus [18] A sand dollar.
Monophoraster M. duboisi [18] A sand dollar.
Neognathostomata Indeterminate An immature specimen.[18] A sea urchin.
Ophiocoma O. sp. [18] A brittle star.
 
Ophiothrix O. sp. [18] A brittle star.
 

Plants

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Plants reported from the Paraná Formation
Genus Species Presence Material Notes Images
Astroniumxylon A. parabalansae Toma Vieja fossil locality.[19] Fossil wood.[19] An anacardiaceaen.
Laurophyllum L. sp. Villa Urquiza.[20] Leaves.[20] A lauraceaen.
Myrciophyllum M. paranaesianum Villa Urquiza.[20] Leaves.[20] A myrtaceaen.
Ocotea O. sp. Villa Urquiza.[21] Leaves.[21] A lauraceaen.
 
Piptadenioxylon P. paraexcelsa Toma Vieja fossil locality.[19] Fossil wood.[19] A mimosoid.
Schinus S. aff. terebinthifolius Villa Urquiza.[20] Leaves.[20] An anacardiaceaen.
 
Solanumxylon S. paranensis Toma Vieja fossil locality.[19][22] Fossil wood.[19] A nightshade.

Correlations

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Laventan

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Laventan correlations in South America
Formation Honda Honda Aisol Cura-Mallín Pisco Ipururo Pebas Capadare Urumaco Inés Paraná Map
Basin VSM Honda San Rafael Caldera Pisco Ucayali Amazon Falcón Venezuela Paraná
Paraná Formation (South America)
Country   Colombia   Bolivia   Argentina   Chile   Peru   Venezuela   Argentina
Boreostemma      
Hapalops      
Miocochilius      
Theosodon      
Xenastrapotherium      
Mylodontidae        
Sparassodonta    
Primates  
Rodents              
Birds      
Terror birds  
Reptiles              
megalodon      
Flora  
Insects  
Environments Fluvial Fluvio-deltaic Fluvio-lacustrine Fluvio-deltaic Fluvial
  Laventan volcanoclastics

  Laventan fauna

  Laventan flora
Volcanic Yes

Huayquerian

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Huayquerian correlations in South America
Formation Cerro Azul Ituzaingó Paraná Camacho Raigón Andalhuala Chiquimil Las Flores Maimará Palo Pebas Muyu Rosa Saldungaray Salicas Urumaco Map
Basin Colorado Paraná Hualfín Tontal Andes Salta Amazon Huasi Altiplano BA Velasco Falcón
Paraná Formation (South America)
Country   Argentina   Uruguay   Argentina   Brazil
  Peru		   Bolivia   Argentina   Venezuela
Cardiatherium        
Lagostomus            
Macroeuphractus      
Proeuphractus        
Pronothrotherium    
Pseudotypotherium      
Thylacosmilus      
Xotodon        
Macraucheniidae            
Primates  
Rodents                      
Reptiles            
Birds        
Terror birds      
Flora    
Environments Aeolian-fluvial Fluvio-deltaic Fluvial Fluvio-lacustrine Fluvial Fluvio-lacustrine Fluvio-deltaic
  Huayquerian volcanoclastics

  Huayquerian fauna

  Huayquerian flora
Volcanic Yes

References

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  1. ^ a b c d e Agnolin, Federico; Lucero, Sergio (December 2004). "Registros de cetáceos del Mioceno Tardío de Entre Ríos, Argentina". Revista de biología marina y oceanografía. 39 (2). doi:10.4067/S0718-19572004000200007. ISSN 0718-1957.
  2. ^ a b c d Pérez, Leandro M.; Cione, Alberto L.; Cozzuol, Mario; Varela, Augusto N. (June 2011). "A Sperm Whale (Cetacea: Physeteroidea) from the Paraná Formation (Late Miocene) of Entre Rís, Argentina. Environment and Taphonomy". Ameghiniana. 48 (4): 648–654. doi:10.5710/AMGH.v48i3(425). ISSN 0002-7014. S2CID 129118367.
  3. ^ a b Noriega, Jorge Ignacio; Cione, Alberto Luis; Aceñolaza, Florencio Gilberto (2007-09-05). "Shark tooth marks on Miocene balaenopterid cetacean bones from Argentina". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 245 (2): 185–192. doi:10.1127/0077-7749/2007/0245-0185. ISSN 0077-7749.
  4. ^ Cozzuol, Mario A. (April 1996). "The record of aquatic mammals in southern South America". Münchner Geowiss. Abh. 30: 321–342.
  5. ^ a b Muizon, Christian de; Hendey, Q. B. (1980). "Late Tertiary seals of the south Atlantic Ocean". Annals of the South African Museum. Annale van die Suid-Afrikaanse Museum. 82: 91–128. ISSN 0303-2515.
  6. ^ de Muizon, Christian; Bond, Mariano (1982). "Le Phocidae (Mammalia) de la Formation Parana (Entre Rios, Argentine)". Bull. Mus. Natn. Hist. Nat., Paris (in French): 165–207.
  7. ^ a b c d e f g h i j Candela, Adriana M.; Bonini, Ricardo A.; Noriega, Jorge I. (November 2012). "First continental vertebrates from the marine Paraná Formation (Late Miocene, Mesopotamia, Argentina): Chronology, biogeography, and paleoenvironments". Geobios. 45 (6): 515–526. Bibcode:2012Geobi..45..515C. doi:10.1016/j.geobios.2012.05.003. S2CID 128431992.
  8. ^ a b c d Vélez-Juarbe, Jorge; Noriega, Jorge I.; Ferrero, Brenda S. (2012). "FOSSIL DUGONGIDAE (MAMMALIA, SIRENIA) FROM THE PARANÁ FORMATION (LATE MIOCENE) OF ENTRE RÍOS PROVINCE, ARGENTINA". Ameghiniana. 49 (4): 585–593. doi:10.5710/AMGH.27.6.2012.568. hdl:11336/72358. ISSN 0002-7014. S2CID 56435219.
  9. ^ a b Noriega, Jorge I.; Cenizo, Marcos; Brandoni, Diego; Pérez, Leandro M.; Tineo, David E.; Diederle, Juan M.; Bona, Paula (2023-05-09). "A new pelican (Aves: Pelecanidae) from the Upper Miocene of Argentina: new clues about the origin of the New World lineages". Journal of Vertebrate Paleontology. 42 (5). doi:10.1080/02724634.2023.2202702. ISSN 0272-4634. S2CID 258605013.
  10. ^ a b c d e f g h i j k l m n o p q Woodward, Arthur Smith (1900). "I.—On some fish-remains from the Parana formation, Argentine Republic". The Annals and Magazine of Natural History; Zoology, Botany, and Geology. 6 (31): 1–7. doi:10.1080/00222930008678334. ISSN 0374-5481.
  11. ^ a b "Tetraodontiformes (Osteichthyes, Actinopterygii) from the Miocene of Argentina: with the southernmost record of fossil Tetraodontidae". Comptes Rendus Palevol. 20 (27): 585–596. 2021.
  12. ^ "Squalus obliquidens | Shark-References". shark-references.com. Retrieved 2023-05-21.
  13. ^ a b Cione, Alberto Luis; Cabrera, Daniel Alfredo; Barla, María Julia (2012-03-01). "Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic". Geobios. 45 (2): 167–172. Bibcode:2012Geobi..45..167C. doi:10.1016/j.geobios.2011.06.002. hdl:11336/139163. ISSN 0016-6995.
  14. ^ a b c Cione, Alberto Luis; Mennucci, Jorge; P&eacute, Leandro; rez; Marí Barla, a Julia (2008-01-01). "Megascyliorhinus trelewensis (Neoselachii) in the Middle-Upper Miocene of Parana, Central Eastern Argentina". Instituto Superior de Correlacion Geologica, Serie Miscelanea. 17 (1): 41–49.
  15. ^ a b c Martín Pérez, Leandro; Griffin, Miguel; Iturerría, Santiago Genta (June 2011). "Pectínidos de la Formación Paraná (Mioceno) Entre Ríos, Argentina". Serie Correlación Geológica (in Spanish). 27 (1): 66–75.
  16. ^ a b c d Péreza, Leandro M.; Iturrería, Santiago F. Genta; Griffini, Miguel (October 2010). "Paleoecological and Paleobiogeographic Significance of Two New Species of Bivalves in the Paraná Formation (Late Miocene) of Entre Ríos Province, Argentina". Malacologia. 53 (1): 61–76. doi:10.4002/040.053.0104. ISSN 0076-2997. S2CID 85017087.
  17. ^ a b c d e f g h i j k l Muravchik, Martín; Griffin, Miguel (2004). "Bryozoans from the Paraná Formation (Miocene), in Entre Ríos province, Argentina" (PDF). Ameghiniana (in Spanish). 41 (1): 3–12. ISSN 1851-8044.
  18. ^ a b c d e f g Martínez, Sergio; Pérez, Leandro M.; del Río, Claudia J. (2021-10-02). "Echinoderms from the upper Miocene Paraná Formation of Argentina". Alcheringa: An Australasian Journal of Palaeontology. 45 (4): 435–442. doi:10.1080/03115518.2021.1985609. ISSN 0311-5518. S2CID 243479952.
  19. ^ a b c d e f Franco, María Jimena (2008-11-30). "Leños fósiles de la Formación Paraná (Mioceno Medio), Toma Vieja, Paraná, Entre Ríos, Argentina: registro de bosques estacionales mixtos". Ameghiniana. 45 (4): 669–717.
  20. ^ a b c d e f Anzótegui, Luisa M.; Aceñolaza, Pablo G. (2008-05-14). "Macrofloristic assemblage of the Parana Formation (Middle-Upper Miocene) in Entre Rios (Argentina)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 248 (2): 159–170. doi:10.1127/0077-7749/2008/0248-0159. hdl:11336/80848. ISSN 0077-7749.
  21. ^ a b Aceñolaza, Pablo G.; Aceñolaza, Florencio G. (1996). "IMPRONTAS FOLIARES DE UNA LAURACEAE EN LA FORMACIÓN PARANÁ (MIOCENO SUPERIOR), EN VILLA URQUIZA, ENTRE RÍOS" (in Spanish). 33 (2): 155–159. ISSN 1851-8044. {{cite journal}}: Cite journal requires |journal= (help)
  22. ^ Franco, M. Jimena; Brea, Mariana (2023-02-27). "Redescription of Solanumxylon paranensis, Late Miocene of Paraná Formation (Entre Ríos, Argentina): A Reliable Fossil Wood of Solanaceae". Ameghiniana. 60 (1). doi:10.5710/AMGH.13.10.2022.3518. ISSN 0002-7014. S2CID 253105168.
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