Introduction

A variety of algae growing on the sea bed in shallow waters
A variety of microscopic unicellular and colonial freshwater algae

Algae (UK: /ˈælɡ/ AL-ghee, US: /ˈæl/ AL-jee; sg.: alga /ˈælɡə/ AL-gə) is an informal term for any organisms of a large and diverse group of photosynthetic eukaryotes, which include species from multiple distinct clades. Such organisms range from unicellular microalgae such as Chlorella, Prototheca and the diatoms, to multicellular macroalgae such as the giant kelp, a large brown alga which may grow up to 50 metres (160 ft) in length. Most algae are aquatic organisms and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds. In contrast, the most complex freshwater forms are the Charophyta, a division of green algae which includes, for example, Spirogyra and stoneworts. Algae that are carried passively by water are plankton, specifically phytoplankton.

Algae constitute a polyphyletic group since they do not include a common ancestor, and although their chlorophyll-bearing plastids seem to have a single origin (from symbiogenesis with cyanobacteria), they were acquired in different ways. Green algae are a prominent examples of algae that have primary chloroplasts derived from endosymbiont cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from endosymbiotic red algae, which they acquired via phagocytosis. Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction via spores.

Algae lack the various structures that characterize plants (which evolved from freshwater green algae), such as the phyllids (leaf-like structures) and rhizoids of bryophytes ( non-vascular plants), and the roots, leaves and other xylemic/phloemic organs found in tracheophytes (vascular plants). Most algae are autotrophic, although some are mixotrophic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy, myzotrophy or phagotrophy. Some unicellular species of green algae, many golden algae, euglenids, dinoflagellates, and other algae have become heterotrophs (also called colorless or apochlorotic algae), sometimes parasitic, relying entirely on external energy sources and have limited or no photosynthetic apparatus. Some other heterotrophic organisms, such as the apicomplexans, are also derived from cells whose ancestors possessed chlorophyllic plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria that produce oxygen as a byproduct of splitting water molecules, unlike other organisms that conduct anoxygenic photosynthesis such as purple and green sulfur bacteria. Fossilized filamentous algae from the Vindhya basin have been dated to 1.6 to 1.7 billion years ago. (Full article...)

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Lake Erie in October 2011, during an intense cyanobacteria bloom

Microcystins—or cyanoginosins—are a class of toxins produced by certain freshwater cyanobacteria, commonly known as blue-green algae. Over 250 different microcystins have been discovered so far, of which microcystin-LR is the most common. Chemically they are cyclic heptapeptides produced through nonribosomal peptide synthases.

Cyanobacteria can produce microcystins in large quantities during algal blooms which then pose a major threat to drinking and irrigation water supplies, and the environment at large. (Full article...)

Selected algae type

Brown algae (sg.: alga) are a large group of multicellular algae comprising the class Phaeophyceae. They include many seaweeds located in colder waters of the Northern Hemisphere. Brown algae are the major seaweeds of the temperate and polar regions. Many brown algae, such as members of the order Fucales, commonly grow along rocky seashores. Most brown algae live in marine environments, where they play an important role both as food and as a potential habitat. For instance, Macrocystis, a kelp of the order Laminariales, may reach 60 m (200 ft) in length and forms prominent underwater kelp forests that contain a high level of biodiversity. Another example is Sargassum, which creates unique floating mats of seaweed in the tropical waters of the Sargasso Sea that serve as the habitats for many species. Some members of the class, such as kelps, are used by humans as food.

Between 1,500 and 2,000 species of brown algae are known worldwide. Some species, such as Ascophyllum nodosum, have become subjects of extensive research in their own right due to their commercial importance. They also have environmental significance through carbon fixation. (Full article...)

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