Epichrysomallidae after Burks et al. 2022 Epichrysomallinae Hill & Riek, 1967. Type genus: Epichrysomalla Girault, 1915.

Epichrysomallidae is a family of gall-forming wasps associated with fig trees (genus Ficus) - they make galls in figs, or on leaves or twigs. Once considered a subfamily of Pteromalidae (Epichrysomallinae), this group of genera has been elevated to family rank;[1] they are now known to be more closely related to other gall-forming chalcid wasps than to pteromalids.

Description

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Antenna with 10–12 flagellomeres, including a small fourth clavomere. The labrum is flexible, and hidden behind the clypeus. Mandibles have three teeth. Notauli complete. In most genera, all legs have five tarsomeres, except for Odontofroggatia and Josephiella, which have four-segmented tarsi. The stigmal vein of the fore wing arises at a right angle and the postmarginal vein is shorter than the stigmal vein.[1]

Genera

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Acophila, Asycobia, Camarothorax, Epichrysomalla, Eufroggattisca, Herodotia, Josephiella, Lachaisea, Leeuweniella, Meselatus, Neosycophila, Odontofroggatia, Parasycobia, Sycobia, Sycobiomorphella, Sycomacophila, Sycophilodes, Sycophilomorpha, Sycotetra.


Pteromalidae after Burks et al. 2022

Colotrechninae

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  • Amerostenini: Amerostenus, Errolia, Glorimontana, Yrka;
  • Colotrechnini: Baridobius, Bofuria, Bomburia, Cameronella, Colotrechnus, Dipachystigma, Dvalinia, Elachertodomyia, Pachyneuronella, Podivna, Uriellopteromalus, Uzka;
  • Divnini: Divna;
  • Trigonoderopsini: Bugacia, Trigonoderopsis.

Erixestinae

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  • Erixestus.

Miscogastrinae

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  • Diconocarini: Diconocara;
  • Miscogastrini: Collentis, Drailea, Glyphognathus, Lamprotatus, Miscogaster, Neoskeloceras, Paralamprotatus, Seladerma, Sphaeripalpus, Stictomischus, Telepsogina, Thektogaster, Tumor, Xestomnaster;
  • Sphegigastrini: Acroclisis, Ammeia, Andersena, Ardilea, Bairamlia, Bubekia, Bubekiana, Callicarolynia, Callimerismus, Ceratetra, Cryptoprymna, Cyrtogaster, Haliplogeton, Halticoptera, Harrizia, Kazina, Maorita, Mauleus, Merismus, Notoglyptus, Notoprymna, Novitzkyanus, Paracroclisis, Ploskana, Polstonia, Rhicnocoelia, Schimitschekia, Sorosina, Sphegigaster, Syntomopus, Thinodytes, Toxeuma, Tricyclomischus, Trigonogastrella, Vespita.

Ormocerinae

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  • Blascoa, Cecidoxenus, Monazosa, Nodisoplata, Ormocerus.

Pachyneurinae

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  • Acroclisoides, Amblyharma, Austroterobia, Canada, Coruna, Euneura, Fusta, Goidanichium, Golovissima, Inkaka, Metastenus, Nazgulia, Neotoxeumorpha, Oomara, Oricoruna, Ottaria, Pachycrepoideus, Pachyneuron, Parabruchobius, Platecrizotes, Teasienna, Toxeumorpha.

Pteromalinae

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  • Otitesellini: Adiyodiella, Apocrypta, Arachonia, Bouceka, Comptoniella, Crossogaster, Diaziella, Dobunabaa, Eujacobsonia, Grandiana, Grasseiana, Guadalia, Lipothymus, Marginalia, Micranisa, Micrognathophora, Otitesella, Philosycella, Philosycus, Philoverdance, Philotrypesis, Robertsia, Seres, Sycoecus, Sycoryctes, Sycoscapter, Walkerella, Watshamiella;
  • Pteromalini: Ablaxia, Abomalus, Acaenacis, Acroclisella, Acroclisissa, Acroclypa, Acrocormus, Aepocerus, Afropsilocera, Aggelma, Agiommatus, Aiemea, Allocricellius, Alticornis, Amandia, Amblypachus, Amphidocius, Angulifrons, Anisopteromalus, Ankaratrella, Anogmoides, Anogmus, Anorbanus, Apelioma, Apsilocera, Apycnetron, Arachnopteromalus, Arriva, Arthrolytus, Asoka, Atrichomalus, Boharticus, Bonitoa, Boucekina, Brachycaudonia, Bupronotum, Caenacis, Caenocrepis, Calliprymna, Callitula, Canberrana, Capellia, Catolaccus, Cecidolampa, Cecidostiba, Cheiropachus, Chlorocytus, Chrysoglyphe, Coelopisthia, Conigastrus, Conomorium, Cratomus, Critogaster, Cyclogastrella, Cyrtophagoides, Cyrtoptyx, Dasyneurophaga, Delisleia, Delucchia, Dibrachoides, Dibrachys, Diglochis, Dimachus, Dinarmoides, Dinarmolaelaps, Dinarmus, Dineuticida, Dinotiscus, Dinotoides, Diourbelia, Dirhicnus, Doganlaria, Dorcatomophaga, Elderia, Endomychobius, Epanogmus, Epicatolaccus, Epipteromalus, Erdoesina, Erythromalus, Eulonchetron, Eumacepolus, Eurydinota, Eurydinoteloides, Eurydinotomorpha, Euteloida, Ezgia, Fedelia, Ficicola, Fijita, Frena, Gbelcia, Genangula, Globimesosoma, Grissellium, Guancheria, Gugolzia, Guinea, Guolina, Gyrinophagus, Habritella, Habritys, Habromalina, Halomalus, Halticopterella, Halticopteroides, Helocasis, Heterandrium, Heteroprymna, Heteroschema, Hillerita, Hlavka, Hobbya, Holcaeus, Homoporus, Huberina, Hypopteromalus, Ischyroptyx, Isocyrtella, Isocyrtus, Isoplatoides, Jaliscoa, Kaleva, Klabonosa, Kratka, Kukua, Kumarella, Lampoterma, Lariophagus, Laticlypa, Lenka, Leodamus, Leptomeraporus, Licteria, Lomonosoffiella, Lonchetron, Longinucha, Lyrcus, Lysirina, Lyubana, Makaronesa, Mazinawa, Megadicylus, Merallus, Meraporus, Merismoclea, Merismomorpha, Merisus, Mesopolobus, Metacolus, Meximalus, Mimencyrtus, Mirekia, Miristhma, Mokrzeckia, Monoksa, Morodora, Muscidifurax, Nadelaia, Narendrella, Nasonia, Neanica, Nedinotus, Neocatolaccus, Neolyubana, Neopolycystus, Nephelomalus, Nikolskayana, Norbanus, Nuchata, Oaxa, Obalana, Olchon, Oniticellobia, Ottawita, Oxyharma, Oxysychus, Pandelus, Panstenon, Paracarotomus, Paradinarmus, Paraiemea, Paroxyharma, Pegopus, Peridesmia, Perilampidea, Perniphora, Pestra, Pezilepsis, Phaenocytus, Platneptis, Platypteromalus, Procallitula, Propicroscytus, Propodeia, Pseudanogmus, Pseudetroxys, Pseudocatolaccus, Psilocera, Psilonotus, Psychophagoides, Psychophagus, Pterapicus, Pterisemoppa, Pteromalus, Pterosemigastra, Pterosemopsis, Ptinocida, Pycnetron, Quercanus, Rakosina, Raspela, Rhaphitelus, Rhopalicus, Rohatina, Roptrocerus, Sceptrothelys, Schizonotus, Sedma, Sigynia, Sisyridivora, Spaniopus, Sphegigastrella, Sphegipterosema, Sphegipterosemella, Spilomalus, Spintherus, Spodophagus, Staurothyreus, Stenetra, Stenomalina, Stenoselma, Stichocrepis, Stinoplus, Strejcekia, Synedrus, Systellogaster, Szelenyinus, Tachingousa, Tanina, Tanzanicesa, Termolampa, Thureonella, Tomicobia, Toxeumella, Toxeumelloides, Trichargyrus, Trichokaleva, Trichomalopsis, Trichomalus, Tricolas, Trimeromicrus, Trinotiscus, Tritneptis, Trjapitzinia, Trychnosoma, Tsela, Uniclypea, Urolepis, Usubaia, Veltrusia, Vrestovia, Xiphydriophagus, Yanchepia, Yosemitea, Zdenekiana.

Sycophaginae

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  • Anidarnes, Conidarnes, Eukoebelea, Idarnes, Neoeukobelea, Pseudidarnes, Sycidiphaga, Sycophaga.

Trigonoderinae

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  • Erdoesia, Eutelisca, Gastracanthus, Janssoniella, Miscogasteriella, Neolelaps, Ogloblinisca, Platygerrhus, Plutothrix, Trigonoderus.

Incertae sedis (unplaced to subfamily)

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Calolelaps, Hemitrichus, Ksenoplata, Mesolelaps, Stictolelaps, Yusufia.

Colotrechninae

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  • Amerostenini: Amerostenus Girault, Errolia Bouček, Glorimontana Bouček, Yrka Bouček;
  • Colotrechnini: Baridobius Heydon, Bofuria Hedqvist, Bomburia Hedqvist, Cameronella Dalla Torre, Colotrechnus Thomson, Dipachystigma Crawford, Dvalinia Hedqvist, Elachertodomyia Girault, Pachyneuronella Girault, Podivna Bouček, Uriellopteromalus Girault, Uzka Bouček;
  • Divnini: Divna Bouček;
  • Trigonoderopsini: Bugacia Erdős, Trigonoderopsis Girault.

Erixestinae

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  • Erixestus Crawford.

Miscogastrinae

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  • Diconocarini: Diconocara Dzhanokmen;
  • Miscogastrini: Collentis Heydon, Drailea Huang, Glyphognathus Graham, Lamprotatus Westwood, Miscogaster Walker, Neoskeloceras Kamijo, Paralamprotatus Liao, Seladerma Walker, Sphaeripalpus Förster, Stictomischus Thomson, Telepsogina Hedqvist, Thektogaster Delucchi, Tumor Huang, Xestomnaster Delucchi;
  • Sphegigastrini: Acroclisis Förster, Ammeia Delucchi, Andersena Bouček, Ardilea Graham, Bairamlia Waterston, Bubekia Dalla Torre, Bubekiana De Santis, Callicarolynia Heydon, Callimerismus Graham, Ceratetra Dzhanokmen, Cryptoprymna Förster, Cyrtogaster Walker, Haliplogeton De Santis, Halticoptera Spinola, Harrizia Delucchi, Kazina Bouček, Maorita Bouček, Mauleus Graham, Merismus Walker, Notoglyptus Masi, Notoprymna De Santis, Novitzkyanus Bouček, Paracroclisis Girault, Ploskana Bouček, Polstonia Heydon, Rhicnocoelia Graham, Schimitschekia Bouček, Sorosina Dzhanokmen, Sphegigaster Spinola, Syntomopus Walker, Thinodytes Graham, Toxeuma Walker, Tricyclomischus Graham, Trigonogastrella Girault, Vespita Bouček.

Ormocerinae

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  • Blascoa Askew, Cecidoxenus Ashmead, Monazosa Dzhanokmen, Nodisoplata Graham, Ormocerus Walker.

Pachyneurinae

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  • Acroclisoides Girault & Dodd, Amblyharma Huang & Tong, Austroterobia Girault, Canada Koçak & Kemal, Coruna Walker, Euneura Walker, Fusta Xiao & Ye, Goidanichium Bouček, Golovissima Dzhanokmen, Inkaka Girault, Metastenus Walker, Nazgulia Hedqvist, Neotoxeumorpha Narendran, Oomara Delucchi, Oricoruna Bouček, Ottaria Hedqvist, Pachycrepoideus Ashmead, Pachyneuron Walker, Parabruchobius Risbec, Platecrizotes Ferrière, Teasienna Heydon, Toxeumorpha Girault.

Pteromalinae

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  • Otitesellini: Adiyodiella Priyadarsanan, Apocrypta Coquerel, Arachonia Joseph, Bouceka Koçak & Kemal, Comptoniella Wiebes, Crossogaster Mayr, Diaziella Grandi, Dobunabaa Bouček, Eujacobsonia Grandi, Grandiana Wiebes, Grasseiana Abdurahiman & Joseph, Guadalia Wiebes, Lipothymus Grandi, Marginalia Priyadarsanan, Micranisa Walker, Micrognathophora Grandi, Otitesella Westwood, Philosycella Abdurahiman & Joseph, Philosycus Wiebes, Philoverdance Priyadarsanan, Philotrypesis Förster, Robertsia Bouček, Seres Waterston, Sycoecus Waterston, Sycoryctes Mayr, Sycoscapter Saunders, Walkerella Westwood, Watshamiella Wiebes;
  • Pteromalini: Ablaxia Delucchi, Abomalus Bouček, Acaenacis Girault, Acroclisella Girault, Acroclisissa Girault, Acroclypa Bouček, Acrocormus Förster, Aepocerus Mayr, Afropsilocera Bouček, Aggelma Delucchi, Agiommatus Crawford, Aiemea Bouček, Allocricellius Yang, Alticornis Bouček, Amandia Graham, Amblypachus De Santis, Amphidocius Dzhanokmen, Angulifrons Xiao & Huang, Anisopteromalus Ruschka, Ankaratrella Risbec, Anogmoides Askew, Anogmus Förster, Anorbanus Bouček, Apelioma Delucchi, Apsilocera Bouček, Apycnetron Bouček, Arachnopteromalus Gordh, Arriva Bouček, Arthrolytus Thomson, Asoka Bouček, Atrichomalus Graham, Boharticus Grissell, Bonitoa Bouček, Boucekina Szelényi, Brachycaudonia Ashmead, Bupronotum Xiao & Huang, Caenacis Förster, Caenocrepis Thomson, Calliprymna Graham, Callitula Spinola, Canberrana Bouček, Capellia Delucchi, Catolaccus Thomson, Cecidolampa Askew, Cecidostiba Thomson, Cheiropachus Westwood, Chlorocytus Graham, Chrysoglyphe Ashmead, Coelopisthia Förster, Conigastrus Bouček, Conomorium Masi, Cratomus Dalman, Critogaster Mayr, Cyclogastrella Bukovskii, Cyrtophagoides Narendran, Cyrtoptyx Delucchi, Dasyneurophaga Hedqvist, Delisleia Girault, Delucchia Koçak & Kemal, Dibrachoides Kurdjumov, Dibrachys Förster, Diglochis Förster, Dimachus Thomson, Dinarmoides Masi, Dinarmolaelaps Masi, Dinarmus Thomson, Dineuticida Bouček, Dinotiscus Ghesquière, Dinotoides Bouček, Diourbelia Risbec, Dirhicnus Thomson, Doganlaria Koçak & Kemal, Dorcatomophaga Kryger, Elderia Hedqvist, Endomychobius Ashmead, Epanogmus Girault, Epicatolaccus Blanchard, Epipteromalus Ashmead, Erdoesina Graham, Erythromalus Graham, Eulonchetron Graham, Eumacepolus Graham, Eurydinota Förster, Eurydinoteloides Girault, Eurydinotomorpha Girault, Euteloida Bouček, Ezgia Koçak & Kemal, Fedelia Delucchi, Ficicola Heydon, Fijita Bouček, Frena Bouček, Gbelcia Bouček, Genangula Bouček, Globimesosoma Xiao & Hui, Grissellium Bouček, Guancheria Hedqvist, Gugolzia Delucchi & Steffan, Guinea Koçak & Kemal, Guolina Heydon, Gyrinophagus Ruschka, Habritella Girault & Dodd, Habritys Thomson, Habromalina Dzhanokmen, Halomalus Erdős, Halticopterella Girault & Dodd, Halticopteroides Girault, Helocasis Wallace, Heterandrium Mayr, Heteroprymna Graham, Heteroschema Gahan, Hillerita Bouček, Hlavka Bouček, Hobbya Delucchi, Holcaeus Thomson, Homoporus Thomson, Huberina Bouček, Hypopteromalus Ashmead, Ischyroptyx Delucchi, Isocyrtella Risbec, Isocyrtus Walker, Isoplatoides Girault, Jaliscoa Bouček, Kaleva Graham, Klabonosa Bouček, Kratka Bouček, Kukua Bouček, Kumarella Sureshan, Lampoterma Graham, Lariophagus Crawford, Laticlypa Bouček, Lenka Bouček, Leodamus Masi, Leptomeraporus Graham, Licteria Risbec, Lomonosoffiella Girault, Lonchetron Graham, Longinucha Bouček, Lyrcus Walker, Lysirina Heydon, Lyubana Bouček, Makaronesa Graham, Mazinawa Bouček, Megadicylus Girault, Merallus Masi, Meraporus Walker, Merismoclea De Santis, Merismomorpha Girault, Merisus Walker, Mesopolobus Westwood, Metacolus Förster, Meximalus Bouček, Mimencyrtus De Santis, Mirekia Bouček, Miristhma Bouček, Mokrzeckia Mokrzecki, Monoksa Bouček, Morodora Gahan, Muscidifurax Girault & Sanders, Nadelaia Bouček, Narendrella Sureshan, Nasonia Ashmead, Neanica Erdős, Nedinotus Bouček, Neocatolaccus Ashmead, Neolyubana Sureshan, Neopolycystus Girault, Nephelomalus Graham, Nikolskayana Bouček, Norbanus Walker, Nuchata Bouček, Oaxa Bouček, Obalana Bouček, Olchon Tselikh, Oniticellobia Bouček, Ottawita Bouček, Oxyharma Bouček, Oxysychus Delucchi, Pandelus Förster, Panstenon Walker, Paracarotomus Ashmead, Paradinarmus Masi, Paraiemea Sureshan & Narendran, Paroxyharma Huang & Tong, Pegopus Förster, Peridesmia Förster, Perilampidea Crawford, Perniphora Ruschka, Pestra Bouček, Pezilepsis Delucchi, Phaenocytus Graham, Platneptis Bouček, Platypteromalus Bouček, Procallitula De Santis, Propicroscytus Szelényi, Propodeia Bouček, Pseudanogmus Dodd & Girault, Pseudetroxys Masi, Pseudocatolaccus Masi, Psilocera Walker, Psilonotus Walker, Psychophagoides Graham, Psychophagus Mayr, Pterapicus Dzhanokmen, Pterisemoppa Girault, Pteromalus Swederus, Pterosemigastra Girault & Dodd, Pterosemopsis Girault, Ptinocida Bouček, Pycnetron Gahan, Quercanus Heydon, Rakosina Bouček, Raspela Bouček, Rhaphitelus Walker, Rhopalicus Förster, Rohatina Bouček, Roptrocerus Ratzeburg, Sceptrothelys Graham, Schizonotus Ratzeburg, Sedma Bouček, Sigynia Hedqvist, Sisyridivora Gahan, Spaniopus Walker, Sphegigastrella Masi, Sphegipterosema Girault, Sphegipterosemella Girault, Spilomalus Graham, Spintherus Thomson, Spodophagus Delvare & Rasplus, Staurothyreus Graham, Stenetra Masi, Stenomalina Ghesquière, Stenoselma Delucchi, Stichocrepis Förster, Stinoplus Thomson, Strejcekia Bouček, Synedrus Graham, Systellogaster Gahan, Szelenyinus Bouček, Tachingousa Tselikh, Tanina Bouček, Tanzanicesa Koçak & Kemal, Termolampa Bouček, Thureonella Gijswijt, Tomicobia Ashmead, Toxeumella Girault, Toxeumelloides Girault, Trichargyrus Dzhanokmen, Trichokaleva Bouček, Trichomalopsis Crawford, Trichomalus Thomson, Tricolas Bouček, Trimeromicrus Gahan, Trinotiscus Bouček, Tritneptis Girault, Trjapitzinia Dzhanokmen, Trychnosoma Graham, Tsela Bouček, Uniclypea Bouček, Urolepis Walker, Usubaia Kamijo, Veltrusia Bouček, Vrestovia Bouček, Xiphydriophagus Ferrière, Yanchepia Bouček, Yosemitea Bouček, Zdenekiana Huggert.

Sycophaginae

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  • Anidarnes Bouček, Conidarnes Farache & Rasplus, Eukoebelea Ashmead, Idarnes Walker, Neoeukobelea Lal, Farooqi & Husain, Pseudidarnes Girault, Sycidiphaga Liu, Rasplus & Huang, Sycophaga Westwood.

Trigonoderinae

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  • Erdoesia, Eutelisca, Gastracanthus, Janssoniella, Miscogasteriella, Neolelaps, Ogloblinisca, Platygerrhus, Plutothrix, Trigonoderus.

Incertae sedis (unplaced to subfamily)

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Calolelaps Timberlake, Hemitrichus Thomson, Ksenoplata Bouček, Mesolelaps Ashmead, Stictolelaps Timberlake, Yusufia Koçak & Kemal

  1. ^ a b Roger A. Burks; Mircea-Dan Mitroiu; Lucian Fusu; et al. (20 December 2022). "From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)". Journal of Hymenoptera Research. 94: 13–88. doi:10.3897/JHR.94.94263. ISSN 1070-9428. Wikidata Q115923766.{{cite journal}}: CS1 maint: unflagged free DOI (link)