Harmonia (beetle)

Harmonia is a genus of lady beetles belonging to the subfamily Coccinellinae.

Harmonia
Asian lady beetle, Harmonia axyridis
Harmonia testudinaria
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Coccinellidae
Subfamily:	Coccinellinae
Genus:	Harmonia Mulsant, 1846
Synonyms
Ballia Mulsant, 1853 Callineda Crotch, 1871 Leis Mulsant, 1850 Ptychanatis Crotch, 1874 Rhopaloneda Timberlake, 1943 Stictoleis Crotch, 1874

Description

The distinction of the individual species within the genus is often very difficult, and in some cases requires examination of the reproductive system. The coloring and drawing in the genus Harmonia are very variable, not only between species, but also within each species. The basic color of the elytra is usually red to orange with black spots. The patches can be linked together to form black bars. The species can reach a length of 3 to 7 mm. The rear edge of the pronotum is slightly curved. The antennae are a little shorter than the head and are composed of eleven segments. The three last segments are thickened at the tip and form a club.

List of species

• Harmonia axyridis (Pallas, 1771) – Asian lady beetle, originally from Nepal, China, Taiwan, Japan, introduced to Western Europe and America
• Harmonia antipodum (Mulsant, 1848) – antipodean ladybird, endemic to New Zealand
• Harmonia basinotata Bielawski, 1964 – New Guinea
• Harmonia bicolor (Blackburn, 1892) – Australia
• Harmonia conformis (Boisduval, 1835) – common spotted lady beetle, Australia, introduced in New Zealand
• Harmonia dimidiata (Fabricius, 1781) – India, Pakistan, Nepal, Bhutan, China, Taiwan, Japan; introduced in North America
• Harmonia dunlopi (Crotch, 1874) – India, Malaysia, Philippines
• Harmonia eucharis (Mulsant, 1850) – India, Pakistan, Myanmar, Nepal, Himalayan Region, South China
• Harmonia expallida (Weise, 1907) – India
• Harmonia octomaculata (Fabricius, 1781) – India, Pakistan, Nepal, Bangladesh, Sri Lanka, Micronesia, Australia
• Harmonia quadripunctata (Pontoppidan, 1763) – cream-streaked lady beetle, Europe
• Harmonia sedecimnotata (Fabricius, 1801) – India, Nepal, Southeast Asia
• Harmonia testudinaria (Mulsant, 1850) – Indonesia, New Guinea, North Australia
• Harmonia uninotata Bielawski, 1964 – New Guinea
• Harmonia yedoensis (Takizawa, 1917) – Japan

References

• Data related to Harmonia at Wikispecies
• Media related to Harmonia at Wikimedia Commons
• Biolib
• Catalogue of Life
• Fauna Europaea